Bellan, Gerard; Bellan-Santini, Denise; Dauvin, Jean-Claude. (1990). Phénétique et phylogénie des Opheliidae (Annélides Polychètes). Comptes-Rendus hebdomadaires des Séances de l'Académie des Sciences de Paris. 310 (5): 175-181. plate 1.
The phenetic and phylogenetic relationships between 12 genera of the family Opheliidae have been studied to attempt a coherent classification of the family. The results show the existence of three monophyletic groups corresponding to the three sub-families previously defined by Hartmann- Schröder
