Holmquist, Charlotte. (1978). Revision of the genus Peloscolex (Oligochaeta, Tubificidae). 1. Morphological and anatomical scrutiny; with discussion on the generic level. Zoologica scripta. 7:187-208.
Re-examination of 35 species and some “varieties” has led to a division of the whole “Peloscolex” complex of species into eight genera, Peloscolex Leidy, 1879, Embolocephalus Randolph, 1892, Orientodrilus gen.n., Baikalodrilus gen.n., Haber gen.n., Edukemius gen.n., and Tubificoides Lastochkin, 1850, Spirosperma Eisen, foremost 1937. This division is based first and on the construction and histology of the male genital organs, but it is also supported by other characteristics.