basis of record
Llewellyn, L. C. (1966). Pontobdellinae (Piscicolidae : Hirudinea) in the British Museum (Natural History) with a review of the subfamily. Bull. Br. Mus. nat. Hist. (Zool.), 14(7), 389-439. [details] Available for editors [request]
From editor or global species database
Ecology typically permanent parasites on elasmobranchs, marine (Sawyer, 1986, p.669) [details]