Marine Biodiversity and Ecosystem Functioning
EU Network of Excellence

Main Menu

· Home
· Contacts
· Data System
· Documents
· Events Calendar
· Forums
· Job M@RKET
· Links
· MarBEF Open Archive
· Network Description
· Outreach
· Photo Gallery
· Quality Assurance
· Register of Resources
· Research Projects
· Rules and Guidelines
· Training
· Weekly News Bulletin
· Wiki
· Worldconference


Last 3 Job opportunities View latest job opportunities RSS feed

>> Go to Job M@RKET

Last 2 forum posts View MarBEF Forum RSS feed

 Field course on sharks in South Africa (June 2011)
 training opportunity in basic taxonomy

>> Go to forums


Who's Online

Currently 33 guest(s) online
Currently 0 member(s) online





Event Calendar

Year View
Month View
Add an Event
Add a Deadline
Jump to the following view:

Training Course on Copepod Biology and Systematics

Location: University of Bizerta (near Tunis city, Tunesia)
Event Type: Training Course
Event Date: 2005-07-04    
End Date: 2005-07-08

Event Description:
Course Objective

The objective of the course is to provide postgraduate students/postdoctoral fellows with a comprehensive and state-of-the-art introduction to the biology, behaviour and morphology of planktonic copepods, plus training in the necessary technical skills.


The course will be taught in English by six tutors: each a specialist in a different area of copepod biology or systematics. The tutors will be:
  • Professor Geoff Boxshall (The Natural History Museum, London, U.K.)
  • Professor Thomas Kirboe (Danish Institute of Fisheries, Denmark)
  • Professor Carol Lee (University of Wisconsin, U.S.A.)
  • Dr Laurent Seuront (CNRS- Marine Station of Wimereux, France)
  • Dr Sami Souissi (University of Lille, France)
  • Professor Rudi Strickler (University of Wisconsin-Milwaukee, U.S.A.)

All tuition is provided FREE. Students attending the course must pay for their own travel to the venue at the University of Bizerta (near Tunis City), Tunisia. The cost of this will be variable. We assume that all participants will be travelling to Tunisia for the 9th International Conference on Copepoda in Hammamet, commencing on 11th July. Transport from the training course to the conference venue will be arranged on 9th July 2005 and a small charge may be made.

Students must also pay for one week's accommodation and meals. Precise costs are not yet available but we estimate that the cost of accommodation plus foodwill be approximately 400 Euros (about 400 USDollars) for six nights/days. More detailed costings will be provided in due course.

How to Apply

All students must complete a questionnaire in order to apply for one of the limited places in the course. The Questionnaire can be found at

This training workshop will be co-sponsored by MARBEF ( and a proportion of the places will be reserved for MARBEF members, who should also completethe questionnaire. Please send or email your completed Questionnaire to Geoff Boxshall at The Natural History Museum, Cromwell Road, London SW7 5BD, UK( The provisional deadline for receipt of applications is 31st December 2004.

Students will be selected in advance by Dr Sami Soussi and Dr Mohamed Nejib Daly Yahia, in consultation with the course organizer. Contacts for practical questions about theUniversity of Bizerta, the samples to be analysed from the local monitoring site of the university, and other questions should be addressed to Mohamed Nejib Daly Yahia( with a copy to

Contact email: or

Event programme:
The course will last five days and will comprise a mix of lectures, laboratory-based practical sessions and informal, problem-solving discussions. We will be flexible and encourage the development of a hot-house atmosphere through the interactions between highly motivated young researchers and established workers.

1) Introduction to Copepod Form and Functional Morphology. Tutor:Geoff Boxshall

A brief overview of copepod morphology and diversity to indicate the range of body forms and of limb types across the whole of the Copepoda. The basic functional morphology of the reproductive,locomotory and feeding systems will also be introduced and a brief synopsis of the current classification system will be presented. This session will provide participants with the morphological foundation onwhich the genetics, behaviour and biology topics can build.

2) Copepod Development. Tutor: Geoff Boxshall

The basic patterns of copepod development will be reviewed and lecithotrophic versus planktotrophic naupliar strategies will be compared. Special emphasis will be placed on the development ofsensory systems and on the correlation between the changing behavioural priorities and changes in the sensor array. This session will begin to focus attention on interactions with the environment andon the comparative structure and [function] of chemosensory, mechanosensory and visual systems.

3) Copepod Behaviour and Small-scale Phenomena Around Individual Copepods. Tutor: Thomas Kirboe

The behaviour and communication of copepods is constrained by the physical and chemical characteristics of their immediate environment (diffusion of solutes, turbulent fluid motion, and dissipation ofmomentum). We will describe fundamental behavioural phenomena such as mate finding, prey finding, and predator avoidance in the context of the governing small-scale physical processes and examine theimplications for population dynamics. The session will consist of lectures and practicals.

4) Observing Copepod Behaviours. Tutor: Rudi Strickler

Copepods are small and whatever they do, they do it fast. The lecture session will concentrate on viewing video clips of copepod behaviours with commentary on what the clips show and why the clipswere taken. Every participant will receive a DVD with the clips for further study. In the practical session we will construct simple optical set-ups which will allow us to register the rapidly executedbehaviours of copepods. Evaluation of the observations will result in graphs and will demonstrate the full cycle from asking the question, then observing the behaviour, to disseminating the results.

5) Copepod Evolutionary Genetics: Understanding Adaptation of Phenotype. Tutor: Carol Lee

The role of genetics in the adaptation of copepods to different environments will be examined. Some overviews using the species complex "Eurytemora affinis" as a model system will be presented.

6) Analysis of Copepod Behaviour Patterns. Tutor: Laurent Seuront

An introduction to different statistical techniques used to characterize the trajectories (2D and 3D) of copepods will be presented. The focus will be on recent techniques(scaling-independent), i.e., fractal dimension, multi-fractal analysis, and random walk techniques. The diversity of copepod behavioural patterns will also be introduced.

7) Modeling Behaviour and Processes at the Individual Level in Copepods. Tutor: Sami Souissi

An introduction to the key processes to be considered when copepods are modeled at the individual scale. A particular attention will be paid to feeding processes (encounter, attack, capture, ingestion) andalso to physiological processes(development, metabolism,reproduction, mortality, etc.). Emerging techniques of modeling basedon multi-agent systems (i.e., Object-Oriented Modeling) will be usedto illustrate thecourse and during practical classes on computer.Recent platforms (software) designed for end-users will be used here in order to make this course accessible to all participants.

Collaborators and organisers:
Geoff Boxshall and Sami Soussi

Website url:

Registration form:

Registration fee: 

Relevant costs that should be taken by MARBEF:  

Possibility for day-care centres (facilities for children): No

MarBEF supported event: Yes

Event within Framework of MARBEF: WP7 - Training courses

Maximum number of participants: 

External participants (non MARBEF) allowed: Yes

Posted by Ward on 2004-05-06 and approved by herman