MarBEF Data System - ERMS - Amphioplus Verrill, 1899

WoRMS taxon details

Amphioplus Verrill, 1899

AphiaID

123611 (urn:lsid:marinespecies.org:taxname:123611)

Classification

Status

accepted

Rank

Genus

Type taxon

Amphiura tumida Lyman, 1878 accepted as Amphioplus tumidus (Lyman, 1878)

Parent

Amphiuridae Ljungman, 1867

Synonymised names

Ailsaria Fell, 1962 · unaccepted (synonym)

Amphiura (Amphioplus) · unaccepted

Direct children (200)

Ordering

Alphabetically
By status

Children Display

direct main ranks all

Subgenus Amphioplus (Amphichilus) Matsumoto, 1917
1. Species Amphioplus (Amphichilus) impressus (Ljungman, 1867)
2. Species Amphioplus (Amphichilus) ochroleuca (Brock, 1888)
3. Species Amphioplus (Amphichilus) trichoides (Matsumoto, 1917)
4. Species Amphioplus (Amphichilus) cesareus (Koehler, 1905) accepted as Amphioplus (Amphichilus) impressus (Ljungman, 1867) (synonymized by A.M. Clark (1970))
5. Species Amphioplus (Amphichilus) daleus (Lyman, 1882) accepted as Silax daleus (Lyman, 1879) (transferred to Unioplus by A.M. Clark (1970), to Silax by Stöhr & O'Hara (2021))
6. Species Amphioplus (Amphichilus) intermedius (Koehler, 1905) accepted as Amphioplus (Amphioplus) intermedius (Koehler, 1905)
Subgenus Amphioplus (Amphioplus) Verrill, 1899
1. Species Amphioplus (Amphioplus) abditus (Verrill, 1871)
2. Species Amphioplus (Amphioplus) aciculatus Mortensen, 1936
3. Species Amphioplus (Amphioplus) ancistrotus Clark, 1911
4. Species Amphioplus (Amphioplus) basilicus (Koehler, 1907)
5. Species Amphioplus (Amphioplus) brasiliensis Tommasi, 1970
6. Species Amphioplus (Amphioplus) debilis (Koehler, 1904)
7. Species Amphioplus (Amphioplus) didymus Clark, 1938
8. Species Amphioplus (Amphioplus) firma (Koehler, 1904)
9. Species Amphioplus (Amphioplus) guangdongensis Liao, 2004
10. Species Amphioplus (Amphioplus) hendleri Liao, 2004
11. Species Amphioplus (Amphioplus) hexabrachiatus Stöhr, 2003
12. Species Amphioplus (Amphioplus) intermedius (Koehler, 1905)
13. Species Amphioplus (Amphioplus) lobata (Ljungman, 1867)
14. Species Amphioplus (Amphioplus) lucidus Koehler, 1922
15. Species Amphioplus (Amphioplus) macilentus (Verrill, 1882)
16. Species Amphioplus (Amphioplus) macraspis (H.L. Clark, 1911)
17. Species Amphioplus (Amphioplus) magellanicus (Mortensen, 1936)
18. Species Amphioplus (Amphioplus) parviclypeus H.L. Clark, 1915
19. Species Amphioplus (Amphioplus) parvitus Cherbonnier & Guille, 1978
20. Species Amphioplus (Amphioplus) pectinatus Mortensen, 1933
21. Species Amphioplus (Amphioplus) personatus (Koehler, 1905)
22. Species Amphioplus (Amphioplus) psilochora (H.L. Clark, 1911)
23. Species Amphioplus (Amphioplus) qingdaoensis Liao, 2004
24. Species Amphioplus (Amphioplus) rhadinobrachius Clark, 1911
25. Species Amphioplus (Amphioplus) sepultus Hendler, 1995
26. Species Amphioplus (Amphioplus) sinicus Liao, 2004
27. Species Amphioplus (Amphioplus) spinosus Cherbonnier & Guille, 1978
28. Species Amphioplus (Amphioplus) stenaspis H.L. Clark, 1938
29. Species Amphioplus (Amphioplus) stewartensis (Mortensen, 1924)
30. Species Amphioplus (Amphioplus) strongyloplax (H.L. Clark, 1911)
31. Species Amphioplus (Amphioplus) tessellata (Koehler, 1904)
32. Species Amphioplus (Amphioplus) thomassini Cherbonnier & Guille, 1978
33. Species Amphioplus (Amphioplus) titubantius Cherbonnier & Guille, 1978
34. Species Amphioplus (Amphioplus) ctenacantha Baker, 1977 accepted as Amphiura ctenacantha (Baker, 1977) (unaccepted > superseded combination, transferred by O'Hara & Thuy, 2022)
35. Species Amphioplus (Amphioplus) furcatus Mortensen, 1933 accepted as Amphioplus (Lymanella) furcatus Mortensen, 1933 (transferred to subgenus Lymanella)
36. Species Amphioplus (Amphioplus) impressa (Ljungman, 1867) accepted as Amphioplus (Amphichilus) impressus (Ljungman, 1867) (transferred to subgenus Amphichilus by Liao & Clark (1995))
37. Species Amphioplus (Amphioplus) impressus (Ljungman, 1867) accepted as Amphioplus (Amphichilus) impressus (Ljungman, 1867) (transferred to subgenus Amphichilus by Liao & Clark (1995))
38. Species Amphioplus (Amphioplus) longirima Fell, 1952 accepted as Amphiura latisquama Mortensen, 1924 (synonymized by Mills & O'Hara (2013))
39. Species Amphioplus (Amphioplus) magnificus (Koehler, 1907) accepted as Silax magnificus (Koehler, 1907) (transferred to Silax by Stöhr % O'Hara (2021))
40. Species Amphioplus (Amphioplus) cyrtacanthus H.L. Clark, 1915 represented as Amphioplus cyrtacanthus H.L. Clark, 1915
Subgenus Amphioplus (Lymanella) A.M. Clark, 1970
1. Species Amphioplus (Lymanella) andreae (Lütken, 1872)
2. Species Amphioplus (Lymanella) depressus (Ljungman, 1867)
3. Species Amphioplus (Lymanella) furcatus Mortensen, 1933
4. Species Amphioplus (Lymanella) hastatus (Ljungman, 1867)
5. Species Amphioplus (Lymanella) integer (Ljungman, 1867)
6. Species Amphioplus (Lymanella) japonicus (Matsumoto, 1915)
7. Species Amphioplus (Lymanella) laevis (Lyman, 1874)
8. Species Amphioplus (Lymanella) peresi Cherbonnier & Guille, 1978
9. Species Amphioplus (Lymanella) potens Koehler, 1930
10. Species Amphioplus (Lymanella) spinulosus (Koehler, 1904)
11. Species Amphioplus (Lymanella) megapomus H.L. Clark, 1911 accepted as Amphioplus (Lymanella) laevis (Lyman, 1874) (synonymized by Liao & Clark (1995))
12. Species Amphioplus (Lymanella) gibbosus Ljungman, 1867 (uncertain > nomen dubium, acc. to A.M. Clark (1970) unrecognizable)
Subgenus Amphioplus (Unioplus) Fell, 1962
1. Species Amphioplus (Unioplus) caulleryi (Koehler, 1897)
2. Species Amphioplus (Unioplus) conditus (Koehler, 1905)
3. Species Amphioplus (Unioplus) confinis (Koehler, 1904)
4. Species Amphioplus (Unioplus) falcatus Mortensen, 1933
5. Species Amphioplus (Unioplus) gentilis (Koehler, 1904)
6. Species Amphioplus (Unioplus) glauca (Lyman, 1879)
7. Species Amphioplus (Unioplus) grata (Koehler, 1904)
8. Species Amphioplus (Unioplus) incisus (Lyman, 1883)
9. Species Amphioplus (Unioplus) pegasus Baker, 1977
10. Species Amphioplus (Unioplus) repositus (Koehler, 1905)
11. Species Amphioplus (Unioplus) servatus (Koehler, 1904)
12. Species Amphioplus (Unioplus) cernuus (Lyman, 1879) accepted as Silax cernuus (Lyman, 1879) (transferred to Silax by Stöhr & O'Hara (2021))
13. Species Amphioplus (Unioplus) cipus Baker, 1977 accepted as Amphiura cipa (Baker, 1977) (unaccepted > superseded combination, transferred by O'Hara & Thuy, 2022)
14. Species Amphioplus (Unioplus) daleus (Lyman, 1879) accepted as Silax daleus (Lyman, 1879) (transferred to Silax by Stöhr & O'Hara (2021))
15. Species Amphioplus (Unioplus) patulus (Lyman, 1879) accepted as Silax patulus (Lyman, 1879) (transferred to Silax by Stöhr & O'Hara (2021))
16. Species Amphioplus (Unioplus) reposita (Koehler, 1905) accepted as Amphioplus (Unioplus) repositus (Koehler, 1905)
Species Amphioplus affinis (Studer, 1885)
Species Amphioplus albidus (Ljungman, 1867)
Species Amphioplus archeri A.M. Clark, 1955
Species Amphioplus asterictus H.L. Clark, 1915
Species Amphioplus aurensis A.M. Clark, 1955
Species Amphioplus brachiostictus Tortonese, 1949
Species Amphioplus caelatus Ely, 1942
Species Amphioplus camamuensis Manso, 2004
Species Amphioplus capax (Koehler, 1905)
Species Amphioplus causatus (Koehler, 1905)
Species Amphioplus cincta (Koehler, 1914)
Species Amphioplus conductus Koehler, 1922
Species Amphioplus congensis (Studer, 1882)
Species Amphioplus coniortodes H.L. Clark, 1918
Species Amphioplus cuneatus (Lyman, 1878)
Species Amphioplus cyrtacanthus H.L. Clark, 1915
Species Amphioplus diacritus Murakami, 1943
Species Amphioplus dispar (Koehler, 1897)
Species Amphioplus echinulatus Mortensen, 1940
Species Amphioplus euryaspis (H.L. Clark, 1911)
Species Amphioplus exsecratus (Koehler, 1905)
Species Amphioplus famula (Koehler, 1910)
Species Amphioplus gravelyi James, 1970
Species Amphioplus heptagonus Cherbonnier & Guille, 1978
Species Amphioplus iuxtus Murakami, 1943
Species Amphioplus legatus Koehler, 1922
Species Amphioplus longuscutum Cherbonnier & Guille, 1978
Species Amphioplus lucyae Tommasi, 1971
Species Amphioplus margueritae Cherbonnier & Guille, 1978
Species Amphioplus mathildae Tommasi & Abreu, 1974
Species Amphioplus minutus Tortonese, 1980
Species Amphioplus modestus (Koehler, 1897)
Species Amphioplus occidentalis Koehler, 1914
Species Amphioplus peregrinator (Koehler, 1912)
Species Amphioplus philohelminthius Ziesenhenne, 1940
Species Amphioplus platyacanthus Murakami, 1943
Species Amphioplus polymorphus Cherbonnier, 1972
Species Amphioplus refectus (Koehler, 1905)
Species Amphioplus seminudus Mortensen, 1940
Species Amphioplus signalis Koehler, 1930
Species Amphioplus somaliensis Tortonese, 1980
Species Amphioplus suspectus Madsen, 1970
Species Amphioplus textilis (Koehler, 1907)
Species Amphioplus thrombodes H.L. Clark, 1918
Species Amphioplus timsae Mortensen, 1926
Species Amphioplus tumidus (Lyman, 1878)
Species Amphioplus abdita (Verrill, 1871) accepted as Amphioplus (Amphioplus) abditus (Verrill, 1871) (spelling error)
Species Amphioplus acanthinus H.L. Clark, 1911 accepted as Dougaloplus acanthinus (H.L. Clark, 1911)
Species Amphioplus acutus Mortensen, 1936 accepted as Amphiura acuta (Mortensen, 1936) (unaccepted > superseded combination, transferred by O'Hara & Thuy, 2022)
Species Amphioplus agassizii Verrill, 1899 accepted as Ophiochiton agassizii (Verrill, 1899) (transferred by A.M. Clark (1970))
Species Amphioplus ailsaclarki Cherbonnier, 1957 accepted as Amphioplus congensis (Studer, 1882)
Species Amphioplus articulatus Mortensen, 1936 accepted as Amphioplus (Amphioplus) aciculatus Mortensen, 1936 (misspelling in Clark (1970), not present in Mortensen (1936))
Species Amphioplus bernasconiae Tommasi, 1968 accepted as Amphioplus albidus (Ljungman, 1867)
Species Amphioplus bocki Koehler, 1927 accepted as Amphioplus (Lymanella) laevis (Lyman, 1874)
Species Amphioplus cinctus (Koehler, 1914) accepted as Amphioplus cincta (Koehler, 1914)
Species Amphioplus cipus Baker, 1977 accepted as Amphiura cipa (Baker, 1977) (unaccepted > superseded combination, transferred by O'Hara & Thuy, 2022)
Species Amphioplus consors (Koehler, 1908) accepted as Silax consors (Koehler, 1907) (transferred to Silax by Stöhr & O'Hara (2021))
Species Amphioplus ctenacantha Baker, 1977 accepted as Amphiura ctenacantha (Baker, 1977) (unaccepted > superseded combination, transferred by O'Hara & Thuy, 2022)
Species Amphioplus cythera A.H. Clark, 1949 accepted as Histampica cythera (A.H. Clark, 1949) (synonym)
Species Amphioplus dalea (Lyman, 1879) accepted as Amphioplus daleus (Lyman, 1879) represented as Amphioplus (Unioplus) daleus (Lyman, 1879) accepted as Silax daleus (Lyman, 1879)
Species Amphioplus depressa (Ljungman, 1867) accepted as Amphioplus (Lymanella) depressus (Ljungman, 1867) (synonym)
Species Amphioplus difficilis (Duncan, 1887) accepted as Ophionephthys difficilis (Duncan, 1887)
Species Amphioplus firmus (Koehler, 1905) accepted as Amphioplus (Amphioplus) firma (Koehler, 1904)
Species Amphioplus formatus (Koehler, 1905) accepted as Ophiodaphne formata (Koehler, 1905)
Species Amphioplus gastracanthus (Lütken & Mortensen, 1899) accepted as Dougaloplus gastracanthus (Lütken & Mortensen, 1899) (transferred by A.M. Clark (1970))
Species Amphioplus glaucus (Lyman, 1879) accepted as Amphioplus (Unioplus) glauca (Lyman, 1879)
Species Amphioplus hexacanthus H.L. Clark, 1911 accepted as Dougaloplus amphacanthus (McClendon, 1909) (synonym)
Species Amphioplus impressa (Ljungman, 1867) accepted as Amphioplus (Amphichilus) impressus (Ljungman, 1867) (misspelling)
Species Amphioplus lobatodes H.L. Clark, 1915 accepted as Amphioplus (Amphioplus) ancistrotus Clark, 1911
Species Amphioplus lobatus (Ljungman, 1867) accepted as Amphioplus (Amphioplus) lobata (Ljungman, 1867)
Species Amphioplus longirima Fell, 1952 accepted as Amphiura latisquama Mortensen, 1924
Species Amphioplus lorioli (Koehler, 1897) accepted as Amphiura (Amphiura) lorioli (Koehler, 1897) (synonym)
Species Amphioplus luctator Koehler, 1922 accepted as Dougaloplus echinatus (Ljungman, 1867)
Species Amphioplus megapomus H.L. Clark, 1911 accepted as Amphioplus (Lymanella) laevis (Lyman, 1874) (synonymized by Liao & Clark (1995))
Species Amphioplus miyadii Murakami, 1943 accepted as Amphioplus (Lymanella) laevis (Lyman, 1874) (synonymized by A.M. Clark (1970))
Species Amphioplus nereis (Lyman, 1883) accepted as Ophiochiton nereis (Lyman, 1883) (transferred by A.M. Clark (1970))
Species Amphioplus relictus (Koehler, 1898) accepted as Amphioplus (Lymanella) depressus (Ljungman, 1867) (synonym)
Species Amphioplus squamata (Delle Chiaje, 1828) accepted as Amphipholis squamata (Delle Chiaje, 1828)
Species Amphioplus stearnsii (Ives, 1891) accepted as Ophiarachnella stearnsii (Ives, 1891)
Species Amphioplus stratus Cherbonnier, 1963 accepted as Amphioplus aciculatus Mortensen, 1936 represented as Amphioplus (Amphioplus) aciculatus Mortensen, 1936 (synonym)
Species Amphioplus tessellatus (Koehler, 1904) accepted as Amphioplus (Amphioplus) tessellata (Koehler, 1904)
Species Amphioplus trepidus (Koehler, 1904) accepted as Ophiomastix flaccida Lyman, 1874 (synonymized by Devaney (1978))
Species Amphioplus velox (Koehler, 1910) accepted as Amphiura (Amphiura) velox Koehler, 1910 (misinterpretation of source)
Species Amphioplus verrilli (Lyman, 1879) accepted as Silax verrilli (Lyman, 1879) (type species of Silax)
Species Amphioplus aciculatus Mortensen, 1936 represented as Amphioplus (Amphioplus) aciculatus Mortensen, 1936 (transferred to subgenus by A.M. Clark (1970))
Species Amphioplus ancistrotus (H.L. Clark, 1911) represented as Amphioplus (Amphioplus) ancistrotus Clark, 1911
Species Amphioplus andreae (Lütken, 1872) represented as Amphioplus (Lymanella) andreae (Lütken, 1872)
Species Amphioplus brasiliensis Tommasi, 1970 represented as Amphioplus (Amphioplus) brasiliensis Tommasi, 1970
Species Amphioplus conditus (Koehler, 1905) represented as Amphioplus (Unioplus) conditus (Koehler, 1905)
Species Amphioplus daleus (Lyman, 1879) represented as Amphioplus (Unioplus) daleus (Lyman, 1879) accepted as Silax daleus (Lyman, 1879) (transferred to Unioplus by A.M. Clark (1970))
Species Amphioplus debilis (Koehler, 1904) represented as Amphioplus (Amphioplus) debilis (Koehler, 1904)
Species Amphioplus depressus (Ljungman, 1867) represented as Amphioplus (Lymanella) depressus (Ljungman, 1867)
Species Amphioplus didymus H.L. Clark, 1938 represented as Amphioplus (Amphioplus) didymus Clark, 1938
Species Amphioplus falcatus Mortensen, 1933 represented as Amphioplus (Unioplus) falcatus Mortensen, 1933
Species Amphioplus furcatus Mortensen, 1933 represented as Amphioplus (Lymanella) furcatus Mortensen, 1933
Species Amphioplus gibbosus (Ljungman, 1867) represented as Amphioplus (Lymanella) gibbosus Ljungman, 1867
Species Amphioplus guangdongensis Liao, 2004 represented as Amphioplus (Amphioplus) guangdongensis Liao, 2004
Species Amphioplus hastatus (Ljungman, 1867) represented as Amphioplus (Lymanella) hastatus (Ljungman, 1867)
Species Amphioplus hendleri Liao, 2004 represented as Amphioplus (Amphioplus) hendleri Liao, 2004
Species Amphioplus hexabrachiatus Stöhr, 2003 represented as Amphioplus (Amphioplus) hexabrachiatus Stöhr, 2003
Species Amphioplus impressus (Ljungman, 1867) represented as Amphioplus (Amphichilus) impressus (Ljungman, 1867) (transferred to subgenus Amphichilus by Liao & Clark (1995))
Species Amphioplus incisus (Lyman, 1883) represented as Amphioplus (Unioplus) incisus (Lyman, 1883)
Species Amphioplus intermedius (Koehler, 1905) represented as Amphioplus (Amphioplus) intermedius (Koehler, 1905)
Species Amphioplus laevis (Lyman, 1874) represented as Amphioplus (Lymanella) laevis (Lyman, 1874)
Species Amphioplus lucidus Koehler, 1922 represented as Amphioplus (Amphioplus) lucidus Koehler, 1922
Species Amphioplus macilentus (Verrill, 1882) represented as Amphioplus (Amphioplus) macilentus (Verrill, 1882)
Species Amphioplus macraspis (H.L. Clark, 1911) represented as Amphioplus (Amphioplus) macraspis (H.L. Clark, 1911)
Species Amphioplus magnificus (Koehler, 1908) represented as Amphioplus (Amphioplus) magnificus (Koehler, 1907) accepted as Silax magnificus (Koehler, 1907)
Species Amphioplus parviclypeus H.L. Clark, 1915 represented as Amphioplus (Amphioplus) parviclypeus H.L. Clark, 1915
Species Amphioplus patulus (Lyman, 1879) represented as Amphioplus (Unioplus) patulus (Lyman, 1879) accepted as Silax patulus (Lyman, 1879)
Species Amphioplus pectinatus Mortensen, 1933 represented as Amphioplus (Amphioplus) pectinatus Mortensen, 1933
Species Amphioplus pegasus Baker, 1977 represented as Amphioplus (Unioplus) pegasus Baker, 1977
Species Amphioplus personatus (Koehler, 1905) represented as Amphioplus (Amphioplus) personatus (Koehler, 1905)
Species Amphioplus potens Koehler, 1930 represented as Amphioplus (Lymanella) potens Koehler, 1930
Species Amphioplus psilochora (H.L. Clark, 1911) represented as Amphioplus (Amphioplus) psilochora (H.L. Clark, 1911)
Species Amphioplus qingdaoensis Liao, 2004 represented as Amphioplus (Amphioplus) qingdaoensis Liao, 2004
Species Amphioplus rhadinobrachius H.L. Clark, 1911 represented as Amphioplus (Amphioplus) rhadinobrachius Clark, 1911
Species Amphioplus sinicus Liao, 2004 represented as Amphioplus (Amphioplus) sinicus Liao, 2004
Species Amphioplus spinosus Cherbonnier & Guille, 1978 represented as Amphioplus (Amphioplus) spinosus Cherbonnier & Guille, 1978
Species Amphioplus stenaspis H.L. Clark, 1938 represented as Amphioplus (Amphioplus) stenaspis H.L. Clark, 1938
Species Amphioplus strongyloplax (H.L. Clark, 1911) represented as Amphioplus (Amphioplus) strongyloplax (H.L. Clark, 1911)
Species Amphioplus thomassini Cherbonnier & Guille, 1978 represented as Amphioplus (Amphioplus) thomassini Cherbonnier & Guille, 1978

Environment

marine, ~~brackish~~, ~~fresh~~, ~~terrestrial~~

Original description

Verrill, A. E. (1899). North American Ophiuroidea. I. Revision of certain families and genera of West Indian ophiurans. Transactions of the Connecticut Academy Arts and Sciences. 10(7), 301-386., available online at https://www.biodiversitylibrary.org/page/13457106#page/331/mode/1up [details]

Taxonomic citation

Stöhr, S.; O’Hara, T.; Thuy, B. (Eds) (2025). World Ophiuroidea Database. Amphioplus Verrill, 1899. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=123611 on 2025-05-31

Taxonomic edit history

Date

action

2004-12-21 15:54:05Z

created

Hansson, Hans

2007-07-12 13:29:01Z

checked

Stöhr, Sabine

Licensing

The webpage text is licensed under a Creative Commons Attribution 4.0 License

[taxonomic tree]

Nomenclature

original description Verrill, A. E. (1899). North American Ophiuroidea. I. Revision of certain families and genera of West Indian ophiurans. Transactions of the Connecticut Academy Arts and Sciences. 10(7), 301-386., available online at https://www.biodiversitylibrary.org/page/13457106#page/331/mode/1up [details]

original description (of Ailsaria Fell, 1962) Fell, H. B. (1962). A revision of the major genera of amphiurid Ophiuroidea. Transactions of the Royal Society of New Zealand, Zoology. 2, 1-26, 1pl. [details]

basis of record Hansson, H.G. (2001). Echinodermata, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels,. 50: pp. 336-351. (look up in IMIS) [details]

Other

context source (PeRMS) Solís-Marín, F. A.; Alvarado, J. J.; Abreu-Pérez, M.; Aguilera, O.; Alió, J.; Bacallado-Aránega, J. J.; Barraza, E.; Benavides-Serrato, M.; Benítez-Villalobos, F.; Betancourt-Fernández, L.; Borges, M.; Brandt, M.; Brogger, M. I.; Borrero-Pérez, G. H.; Buitrón-Sánchez, E.; Campos, L. S.; Cantera, J.; Clemente, S.; Cohen-Renjifo, M.; Coppard, S.; Costa-Lotufo, L. V.; del Valle-García, R.; Díaz, Y.; Díaz de Vivar, M. E.; Díaz-Martínez, J. P.; Durán-González, A.; Epherra, L.; Escolar, M.; Francisco, V.; Freire, C. A.; García-Arrarás, E.; Gil, D. G.; Guarderas, P.; Hadel, V. F.; Hearn, A.; Hernández, J. C.; Hernández-Delgado, E. A.; Herrera-Moreno, A.; Herrero-Pérezrul, M. D.; Hooker, Y.; Honey-Escandón, M. B. I.; Lodeiros, C.; Luzuriaga, M.; Manso, C. L. C.; Martín, A.; Martinez, M. I.; Martínez, S.; Moro-Abad; Mutschke, E.; Navarro, J. C.; Neira, R.; Noriega, N.; Palleiro-Nayar, J. S.; Pérez, A. F.; Pérez-Ruzafa, A.; Prieto-Rios, E.; Reyes, J.; Rodríguez, R.; Rubilar, T.; Sancho-Mejía, T.; Sangil, C.; Silva, J. R. M. C.; Sonnenholzner, J. I.; Ventura, C. R.; Tablado, A.; Tavares, Y.; Tiago, C. G.; Tuya, F.;Williams, S. M. (2013). Appendix. In: J. J. Alvarado & F. A. Solís-Marín (eds), Echinoderm Research and Diversity in Latin America. pp. 471-510. Springer; Berlin & Heidelberg. page(s): 543-654. [details]

additional source Clark, A. M. (1974). Notes on some echinoderms from southern Africa. Bulletin of the British Museum of Natural History (Zoology). 26(6): 423-487, 3 pls, 16 figs, 1 map, 3 tables., available online at https://www.biodiversitylibrary.org/page/2294231 [details]