MarBEF Data System



WoRMS taxon details

Kinorhyncha

101060  (urn:lsid:marinespecies.org:taxname:101060)

Reinhard, 1885
accepted
Phylum

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  1. Class Allomalorhagida
  2. Class Cyclorhagida
  3. Order Cyclorhagae accepted as Cyclorhagida (emendation of name)
  4. Class Homalorhagae accepted as Allomalorhagida (new combination)
  5. Order Homalorhagida accepted as Allomalorhagida (new combination)
marine, brackish
Reinhard, W. (1885). Kinorhyncha (Echinoderes), leur structure anatomique et leur place dans le système. <em>Travaux de la Société des Naturalistes à l'Université Impériale de Kharkow.</em> 19: 205-305, pls. VIII-XII. [in Russian]. [details] OpenAccess publication
Etymology "Reinhard (1885: p. 296, 1887: p. 463) chooses the name “Kinorhyncha” (kinein = Greek for move, rhynchos = Greek for...  
Etymology "Reinhard (1885: p. 296, 1887: p. 463) chooses the name “Kinorhyncha” (kinein = Greek for move, rhynchos = Greek for snout), because he wants to avoid confusion with, e.g., Echinodermata, and because he misidentifies the posterior extensions of the esophagus bulging between the pharynx protractors as brain and consequently the protrusible introvert as protrusible snout of the alimentary canal (Zelinka 1928: p. 38)." [details]

Taxonomy Higgins (1986): species list of Kinorhyncha.  
Taxonomy Higgins (1986): species list of Kinorhyncha. [details]

Taxonomy Higgins, R. P. (1981): The author described the taxonomic ranks within the phylum Kinorhyncha from order to genus level and...  
Taxonomy Higgins, R. P. (1981): The author described the taxonomic ranks within the phylum Kinorhyncha from order to genus level and provided illustrations of the genera. [details]
Neuhaus, B. (2024). World Kinorhyncha Database. Kinorhyncha. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=101060 on 2024-11-21
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2004-12-21 15:54:05Z
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2020-01-20 13:40:57Z
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original description Reinhard, W. (1885). Kinorhyncha (Echinoderes), leur structure anatomique et leur place dans le système. <em>Travaux de la Société des Naturalistes à l'Université Impériale de Kharkow.</em> 19: 205-305, pls. VIII-XII. [in Russian]. [details] OpenAccess publication

taxonomy source Reinhard, W. (1887). Kinorhyncha (Echinoderes), ihr anatomischer Bau und ihre Stellung im System. <em>Zeitschrift für wissenschaftliche Zoologie.</em> 45: 401-467, pls. XX-XXII.
note: Reinhard (1887) represented the German translation of the Russian article of Reinhard (1885). Unfortunately, Reinhard (1887) made only in the title a single, a bit cryptic reference to his earlies stu...    
Reinhard (1887) represented the German translation of the Russian article of Reinhard (1885). Unfortunately, Reinhard (1887) made only in the title a single, a bit cryptic reference to his earlies study from 1885 in Russian, which is the taxonomically relevant article. Therefore, Reinhard (1885) seemed to have escaped the attention of the scientific community for a long time (see also Neuhaus 2013, p. 273).
[details] OpenAccess publication

taxonomy source Malakhov, V. V. (1980). Cephalorhyncha, a new animal phylum uniting Priapulida, Kinorhyncha, Gordiacea, and a system of Aschelminthes worms [in Russian]. <em>Zoologichesky Zhurnal.</em> 59(4): 485-499. [details] Available for editors  PDF available [request]

taxonomy source Adrianov, A. V.; Malakhov, V. V. (1999). Cephalorhyncha of the world ocean. <em>KMK Scientific Press, Moscow.</em> 328 pp. [in Russian and English].
note: The book reviews the research history, anatomy, postembryonic development, biogeography, phylogeny, and systematics at species level of Priapulida, Loricifera, Nematomorpha, and Kinorhyncha in Russian...    
The book reviews the research history, anatomy, postembryonic development, biogeography, phylogeny, and systematics at species level of Priapulida, Loricifera, Nematomorpha, and Kinorhyncha in Russian and the latter taxon also in English. Identification keys at species level are provided. All species are briefly treated and illustrated, 13 new species are described.
[details] Available for editors  PDF available [request]

taxonomy source Sørensen, M. V.; Dal Zotto, M.; Rho, H. S.; Herranz, M.; Sánchez, N.; Pardos, F.; Yamasaki, H. (2015). Phylogeny of Kinorhyncha based on morphology and two molecular loci. <em>PLOS ONE.</em> 10(7): e0133440, 1-33., available online at https://doi.org/10.1371/journal.pone.0133440 [details] Available for editors  PDF available [request]

taxonomy source Sørensen, M. V. (2013). Phylum Kinorhyncha. <em>In</em>: Zhang, Z.-Q. (Ed.) Animal Biodiversity: An outline of higher-level classification and survey of taxonomic richness (Addenda 2013). Zootaxa 3703, 1-82. <em>Zootaxa.</em> 3703(1): 63-66., available online at https://doi.org/10.11646/zootaxa.3703.1.13
note: Outline of classification of Kinorhyncha with orders, families, and genera as well as species numbers of each genus. [details] 

taxonomy source Telford, M. J.; Bourlat, S. J.; Economou, A.; Papillon, D.; Rota-Stabelli, O. (2008). The evolution of the Ecdysozoa. <em>Philosophical Transactions of the Royal Society B: Biological Sciences.</em> 363: 1529-1537., available online at https://doi.org/10.1098/rstb.2007.2243
note: Abstract (shortened): Ecdysozoa is a clade composed of arthropods, tardigrades, onychophorans, nematodes, nematomorphs, priapulids, kinorhynchs and loriciferans. The monophyly of the clade has been ca...    
Abstract (shortened): Ecdysozoa is a clade composed of arthropods, tardigrades, onychophorans, nematodes, nematomorphs, priapulids, kinorhynchs and loriciferans. The monophyly of the clade has been called into question by some workers based on analyses of whole genome datasets. We review the evidence that now conclusively supports the unique origin of these phyla. Relationships within Ecdysozoa are also controversial and we discuss the molecular and morphological evidence for a number of monophyletic groups within this superphylum.
[details] Available for editors  PDF available [request]

taxonomy source Chitwood, B G. (1959). The classification of the phylum Kinorhyncha (Reinhard, 1887) Pearse, 1936. <em>Proceedings of the XVth International Congress of Zoology, London, 16-23 July 1958, Hewer, H. R., & Riley, N. D. (eds), XVth International Congress of Zoology at the Rooms of the Linnean Society of London.</em> 941-943.
note: The article provides a classification of Kinorhyncha. [details] Available for editors  PDF available [request]

taxonomy source Giribet, G.; Edgecombe, G. D. (2017). Current understanding of Ecdysozoa and its internal phylogenetic relationships. <em>Integrative and Comparative Biology.</em> 57(3): 455-466., available online at https://doi.org/10.1093/icb/icx072 [details] 

taxonomy source Laumer, C. E.; Fernández, R.; Lemer, S.; Combosch, D.; Kocot, K. M.; Riesgo, A.; Andrade, S. C. S.; Sterrer, W.; Sørensen, M. V.; Giribet, G. (2019). Revisiting metazoan phylogeny with genomic sampling of all phyla. <em>Proceedings of the Royal Society B: Biological Sciences.</em> 286, 20190831: 1-10., available online at https://doi.org/10.1098/rspb.2019.0831
note: We here present the first attempt to investigate (though not conclusively resolve) animal relationships using genome-scale data from all phyla. Results from the site-heterogeneous CAT þ GTR model rec...    
We here present the first attempt to investigate (though not conclusively resolve) animal relationships using genome-scale data from all phyla. Results from the site-heterogeneous CAT þ GTR model recapitulate many established major clades, and strongly confirm some recent discoveries, such as a monophyletic Lophophorata, and a sister group relationship between Gnathifera and Chaetognatha, raising continued questions on the nature of the spiralian ancestor. We also explore matrix construction with an eye towards testing specific relationships; this approach uniquely recovers support for Panarthropoda, and shows that Lophotrochozoa (a subclade of Spiralia) can be constructed in strongly conflicting ways using different taxon- and/or orthologue sets. Dayhoff-6 recoding sacrifices information, but can also reveal surprising outcomes, e.g. full support for a clade of Lophophorata and Entoprocta þ Cycliophora, a clade of Placozoa þ Cnidaria, and raising support for Ctenophora as sister group to the remaining Metazoa, in a manner dependent on the gene and/or taxon sampling of the matrix in question. Future work should test the hypothesis that the few remaining uncertainties in animal phylogeny might reflect violations of the various stationarity assumptions used in contemporary inference methods.
[details] Available for editors  PDF available [request]

taxonomy source Jeeva, C.; Mohan, P. M.; Ragavan, P.; Muruganantham, V. (2020). A review on taxonomy of phylum Kinorhyncha. <em>Open Journal of Marine Science.</em> 10(4): 260-294., available online at https://doi.org/10.4236/ojms.2020.104020
note: Editor's comment: The authors considered species indeterminata, taxa inquirenda, nomina dubia, and species described from juvenile stages as invalid species, which does not agree with the ICZN. [details] 

taxonomy source Herranz, M.; Stiller, J.; Worsaae, K.; Sørensen, M. V. (2022). Phylogenomic analyses of mud dragons (Kinorhyncha). <em>Molecular Phylogenetics and Evolution.</em> 168, 107375: 1-10., available online at https://doi.org/10.1016/j.ympev.2021.107375 [details] Available for editors  PDF available [request]

taxonomy source Pardos, F.; Cepeda, D. (2024). Kinomenclature: An analysis of scientific names of mud dragons (Kinorhyncha). <em>Zoologischer Anzeiger.</em> 313: 279-288., available online at https://doi.org/10.1016/j.jcz.2024.10.014 [details] Available for editors  PDF available [request]

basis of record Neuhaus, B.; van der Land, J. (2001). Cephalorhyncha (=Loricifera, Priapulida, Kinorhyncha, Nematomorpha), <B><I>in</I></B>: Costello, M.J. <i>et al.</i> (Ed.) (2001). <i>European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels,</i> 50: pp. 159-160 (look up in IMIS) [details] OpenAccess publication

additional source Greeff, R. (1869). Untersuchungen über einige merkwürdige Thiergruppen des Arthropoden- und Wurm-Typus. <em>Archiv für Naturgeschichte.</em> 35(1): 71-121., available online at https://www.biodiversitylibrary.org/page/7058414
note: article covers history, morphology, ecology and phylogeny of Kinorhyncha [details] OpenAccess publication

additional source Zelinka, C. (1908). Zur Anatomie der Echinoderen. <em>Zoologischer Anzeiger.</em> 33 (19/20): 629-647.
note: The article reports about kinorhynch morphology with an intensive discussion of Schepotieff (1908). The author also introduces inadvertently Centropsis parallela, Pycnophyes communis, P. flaveolata, a...    
The article reports about kinorhynch morphology with an intensive discussion of Schepotieff (1908). The author also introduces inadvertently Centropsis parallela, Pycnophyes communis, P. flaveolata, and Trachyemus giganteus as new species.
[details] OpenAccess publication

additional source Zelinka, K. (1928). Monographie der Echinodera. <em>Verlag von Wilhelm Engelmann, Leipzig.</em> 1-396, plates I-XXVII.
note: The book covers almost all aspects of Kinorhyncha, including morphology, reproduction, postembryonic development, phylogeny, parasites, ecology, distribution, and taxonomy. [details] Available for editors  PDF available [request]

additional source Neuhaus, B. (2013). 5. Kinorhyncha (= Echinodera). In: Schmidt-Rhaesa, A. (Ed.), Handbook of Zoology, Gastrotricha, Cycloneuralia and Gnathifera, Volume 1: Nematomorpha, Priapulida, Kinorhyncha, Loricifera. Walter de Gruyter, Berlin,. pp. 181-348. (look up in IMIS), available online at https://doi.org/10.1515/9783110272536.181
note: The book chapter reviews all known aspects of Kinorhyncha like research history, taxonomy, phylogeny, morphology, anatomy, reproduction, locomotion, ecology, development, biogeography, epifauna, and p...    
The book chapter reviews all known aspects of Kinorhyncha like research history, taxonomy, phylogeny, morphology, anatomy, reproduction, locomotion, ecology, development, biogeography, epifauna, and parasites. A key at genus level is also provided.
[details] Available for editors  PDF available [request]

additional source Schepotieff, A. (1907). Die Echinoderiden. <em>Zeitschrift für wissenschaftliche Zoologie.</em> 88(2): 291-326, pls. XVII-XX.
note: The article provides information about history of research and new information about the anatomy, morphology and distribution of several species of Echinoderes. [details] OpenAccess publication

additional source Zelinka, C. (1907). Zur Kenntnis der Echinoderen. <em>Zoologischer Anzeiger.</em> 32(5): 130-136.
note: The article reports about kinorhynch morphology with an intensive discussion of Schepotieff (1907). [details] OpenAccess publication

additional source Higgins, R.P. & H. Thiel. (1988). Introduction to the Study of Meiofauna. <em>Smithsonian Institution Press, Washington, D.C.</em> 488 pp.
note: The book chapter reviews briefly the classification, habitats, collection and further processing of Kinorhyncha. [details] Available for editors  PDF available [request]

additional source Adrianov, A. V. and Malakhov, V. V. (1995). The phylogeny and classification of the phylum Cephalorhyncha. <em>Zoosystematics Rossica.</em> 3(2): 181-201.
note: The article argues about a phylogenetic system of Cephalorhyncha comprising fossila and recent taxa of Priapulida, Kinorhyncha, Loricifera, and Nematomorpha. [details] Available for editors  PDF available [request]

additional source Bütschli, O. (1876). Untersuchungen über freilebende Nematoden und die Gattung Chaetonotus. Zeitschrift für wissenschaftliche Zoologie, 26: 363-413., available online at https://www.biodiversitylibrary.org/itempdf/135000
note: The article discussed the phylogenetic relationsships of Kinorhyncha to other worm groups. [details] 

additional source Reinhard, W. (1883). Zum Bau der Echinoderes. <em>Protokoll der VII. Versammlung russischer Naturforscher und Ärzte. Sitzung der zoologischen Sektion den 22. August 1883, Odessa.</em> Vol. 2A (3): 1-11 [in Russian]. [details] OpenAccess publication

additional source Yamasaki, H.; Hiruta, S. F.; Kajihara, H. (2013). Molecular phylogeny of kinorhynchs. <em>Molecular Phylogenetics and Evolution.</em> 67(2): 303-310., available online at https://doi.org/10.1016/j.ympev.2013.02.016
note: The article represented the first analysis of the phylogenetic relationsships within Kinorhyncha based on molecular data. [details] Available for editors  PDF available [request]

additional source Schepotieff, A. (1908). 3. Zur Kenntnis der Echinoderiden. <em>Zoologischer Anzeiger.</em> 32: 585-589.
note: The article reports about kinorhynch morphology with an intensive discussion of Zelinka (1907). [details] OpenAccess publication

additional source van der Land, J. (ed). (2008). UNESCO-IOC Register of Marine Organisms (URMO). , available online at http://www.marinespecies.org/urmo/ [details] 

additional source Remane, A. (1929). Dritte Klasse des Cladus Nemathelminthes. Kinorhyncha = Echinodera. <em>De Gruyter, Berlin, Handbuch der Zoologie.</em> Band 2, Lieferung 6, 187-248.
note: The book chapter covers almost all aspects of Kinorhyncha, including morphology, reproduction, postembryonic development, phylogeny, parasites, ecology, distribution, and taxonomy. [details] Available for editors  PDF available [request]

additional source Kristensen, R. M. (2002). Kinorhyncha. <em>Encyclopedia of Life Sciences. John Wley & Sons, Ltd.</em> 1-3., available online at https://doi.org/10.1038/npg.els.0001590
note: Short review of Kinorhyncha. [details] Available for editors  PDF available [request]

additional source González-Casarrubios, A.; Sánchez, N. (2024). Bibliometrics of Scalidophora (Animalia: Ecdysozoa): Evolution of knowledge, research topics and scientific collaborations in three distinct animal phyla. <em>Zoologischer Anzeiger.</em> 313: 332-344., available online at https://doi.org/10.1016/j.jcz.2024.10.015 [details] Available for editors  PDF available [request]

additional source Neuhaus, B. (2022). How repeatable are scientific studies of Kinorhyncha? An analysis of specimen-based location and deposition data in WoRMS from 1863 to 2020. <em>Zoologischer Anzeiger.</em> 301: 163-173. (look up in IMIS), available online at https://doi.org/10.1016/j.jcz.2022.08.006 [details] Available for editors  PDF available [request]

additional source Adrianov, A. V.; Malakhov, V. V. (1994). Kinorhyncha: Structure, development, phylogeny and taxonomy. <em>Nauka Publishing, Moscow.</em> 1–262. [in Russian].
note: The book describes light microscopy, TEM, SEM, postembryonic development, phylogeny and taxonomy of Kinorhyncha and provides identification keys at species llevel of species-rich taxa. [details] Available for editors  PDF available [request]

additional source de Beauchamp, P. (1965). Classe des kinorhynques. Echinodères (Dujardin 1851)—Kinorhyncha (Reinhard 1887). <em>In: Grassé, P.-P. (ed.), Traité de Zoologie. Anatomie, systématique, biologie. Masson et Cie Éditeurs Libraires de l'Académie de Médecine, Paris.</em> Vol. IV Némathelminthes (nématodes, gordiacés, rotifères, gastrotriches, kinorhynques), fascicule 3: 1407–1419.
note: The book chapter covers almost all aspects of Kinorhyncha, including morphology, reproduction, postembryonic development, phylogeny, distribution, and taxonomy. [details] Available for editors  PDF available [request]

additional source Adrianov, A. V.; Malakhov, V. V. (1996). The phylogeny and classification of the class Kinorhyncha. <em>Zoosystematics Rossica.</em> 4 (1): 23-44.
note: The article provides a phylogenetic analysis and a classification of Kinorhyncha. [details] Available for editors  PDF available [request]

additional source Remane, A. (1936). Aschelminthes. Kinorhyncha (= Echinodera). In: Remane, A., Gastrotricha und Kinorhyncha, Dr. H. G. Bronn's Klassen und Ordnungen des Tierreichs. Akademische Verlagsgesellschaft, Leipzig. 4. Band: Vermes, II. Abteilung: Aschelminthes, Trochelminthes, 1. Buch, 2.Teil: Gastrotricha und Kinorhyncha, pp.243-372, Appendix pp. 373–385.
note: The book chapter covers almost all aspects of Kinorhyncha, including morphology, reproduction, postembryonic development, phylogeny, parasites, ecology, distribution, and taxonomy. [details] Available for editors  PDF available [request]

additional source Needham, A. E. (1989). 10. Kinorhyncha. In: Adiyodi, K.G. & Adiyodi, R.G. (eds). <em>Reproductive biology of invertebrates.</em> Vol IV, Part A: Fertilization, development, and parental care. John Wiley & Sons, Chichester, pp. 207–217.
note: The book chapter reviews aspects of the reproductive biology of Kinorhyncha. [details] Available for editors  PDF available [request]

additional source Adrianov, A. V.; Malakhov, V. V. (1995). Synopsis of the system of the phylum Cephalorhyncha. <em>Russian Journal of Marine Biology.</em> 21 (2): 89-96.
note: The article suggests a classification of Cephalorhyncha comprising fossila and recent taxa of Priapulida, Kinorhyncha, Loricifera, and Nematomorpha. [details] Available for editors  PDF available [request]

status source Ruggiero, M.A., D.P. Gordon, T.M. Orrell, N. Bailly, T. Bourgoin, R.C. Brusca, T. Cavalier-Smith, M.D. Guiry & P.M. Kirk. (2015). A Higher Level Classification of All Living Organisms. <em>PLOS ONE.</em> 10(4):1-60. (look up in IMIS), available online at https://doi.org/10.1371/journal.pone.0119248 [details] OpenAccess publication

ecology source Cepeda, D.; Pardos, F.; Sánchez, N. (2021). From biggest to smallest mud dragons: size-latitude trends in a group of meiobenthic animals worldwide. <em>Organisms Diversity & Evolution.</em> 21(1): 43-58., available online at https://doi.org/10.1007/s13127-020-00471-y [details] Available for editors  PDF available [request]

identification resource Boykin, J. C. (1974). Phylum Kinorhyncha. In: E.N. Kozloff (ed.), Keys to the marine invertebrates of Puget Sound and the San Juan Archipelago, and adjacent regions. VII + 266 pp. University of Washington Press, Seattle. pp. 39-41. [details] Available for editors  PDF available [request]

identification resource Adrianov, A. V.; Maiorova, A. (2020). Chapter 13 Kinorhyncha. A review on biogeography of the deep-sea Kinorhyncha (phylum Cephalorhyncha Malakhov, 1980) along the NW Pacific. In: Saeedi, H. & Brandt, A. (eds), Biogeographic atlas of the deep NW Pacific fauna. 251-266., available online at https://doi.org/10.3897/ab.e51315
note: The authors summarize information about deep-sea Kinorhyncha from the Kuril-Kamchatka trench and comment about their diagnoses and biogeography. [details] 

identification resource Chatterjee, T. (2019). A review note on research studies of Kinorhyncha from India. <em>Cahiers de Biologie Marine.</em> 60: 183-188., available online at http://application.sb-roscoff.fr/cbm/article.htm?execution=e7s1&_eventId=downloadpdf&articleId=41079 [details] Available for editors  PDF available [request]

identification resource Sørensen, M. V.; Pardos, F. (2008). Kinorhynch systematics and biology – an introduction to the study of kinorhynchs, inclusive identification keys to the genera. <em>Meiofauna Marina.</em> 16: 21-73.
note: The authors describe and illustrate the morphological characters necessary for identification of Kinorhyncha at genus level, habitats, collection, sample processing including mounting for light micros...    
The authors describe and illustrate the morphological characters necessary for identification of Kinorhyncha at genus level, habitats, collection, sample processing including mounting for light microscopy, glossary of terms, and provide an illustrated identification key at genus level.
[details] Available for editors  PDF available [request]

identification resource Bacescu, M. (1968). Class Kinorhyncha. Determination of the fauna of the Black and Azov Seas. 3rd edition. <em>Naukova Dumka.</em> 1, Free-living invertebrates: 237-250 [in Russian].
note: The authors provide a rough key for identification of Kinorhyncha from the Black Sea and Azov Sea. [details] Available for editors  PDF available [request]

identification resource Dal Zotto, M.; Todaro, M. A. (2016). Kinorhyncha from Italy, a revision of the current checklist and an account of the recent investigations. <em>Zoologischer Anzeiger - A Journal of Comparative Zoology.</em> 265: 90-107., available online at https://doi.org/10.1016/j.jcz.2016.01.004 [details] Available for editors  PDF available [request]

identification resource Sørensen, M. V.; Pardos, F. (2020). Kinorhyncha. In: Schmidt-Rhaesa, A. (Ed.), Guide to the identification of marine meiofauna. Verlag Dr. Friedrich Pfeil, Munich. 391-414.
note: The authors describe and illustrate the morphological characters necessary for identification of Kinorhyncha at genus level, habitats, extraction, and provide an illustrated identification key at genu...    
The authors describe and illustrate the morphological characters necessary for identification of Kinorhyncha at genus level, habitats, extraction, and provide an illustrated identification key at genus level.
[details] Available for editors  PDF available [request]

identification resource Adrianov, A. V. (2011). Phylum Cephalorhyncha, class Kinorhyncha. In: Buzhinskaja, G.N. (ed.), Illustrated key to the invertebrates of Eurasien Arctic Seas and adjacent deep waters, Vol. 2 Nemertea, Cephalorhyncha, Oligochaeta, Hirudinida, Pogonophora, Echiura, Sipuncula, Phoronida, and Brachiopoda. <em>Alaska Sea Grant, University of Alaska Fairbanks, Fairbanks.</em> pp. 41-49.
note: The author provides an illustrated key for the identification of Eurasien Arctic Kinorhyncha at species level. [details] Available for editors  PDF available [request]

identification resource Bacescu, M. (1968). Class Kinorhyncha. Determination of the fauna of the Black and Azov Seas. 3rd edition. [2 English translations]. <em>Naukova Dumka.</em> 1, Free-living invertebrates: 237-250.
note: The authors provide a rough key for identification of Kinorhyncha from the Black Sea and Azov Sea. [details] Available for editors  PDF available [request]

identification resource Neuhaus, B. (2004). 19. Kinorhyncha. In: Hofrichter, R. (ed.), Das Mittelmeer. Fauna, Flora, Ökologie. Vol. II/2 Bestimmungsführer, pp. 646-653. Spektrum Akademischer Verlag. [details] Available for editors  PDF available [request]

identification resource Yamasaki, H. (2017). Diversity of Kinorhyncha in Japan and phylogenetic relationships of the phylum. In: Motokawa, M. & Kajihara, H. (eds), Species diversity of animals in Japan, diversity and commonality in animals. Springer Press, Tokyo. pp. 543-563., available online at https://doi.org/10.1007/978-4-431-56432-4_21
note: The book chapter reviews the extraction of specimens, the research history of Kinorhyncha in Japan, and the phylogeny of the group, provides a species record list and and multi-entry identification gu...    
The book chapter reviews the extraction of specimens, the research history of Kinorhyncha in Japan, and the phylogeny of the group, provides a species record list and and multi-entry identification guide to genus level.
[details] Available for editors  PDF available [request]

identification resource Remane, A. (1928). VII. d2 Kinorhyncha. In: Grimpe, G. & Wagler, E. (eds), Tierwelt der Nord- und Ostsee. Akademische Verlagsgesellschaft Becker & Erler, Leipzig. VIId5: 57-84.
note: The book chapter provides general information about the study of Kinorhyncha, their morphology, postembryonic development, distribution, ecology, and phylogenetic relationships, as well as an identifi...    
The book chapter provides general information about the study of Kinorhyncha, their morphology, postembryonic development, distribution, ecology, and phylogenetic relationships, as well as an identification key at genus level and sometimes at species level.
[details] Available for editors  PDF available [request]

biology source Neuhaus, B. (2004). 19. Kinorhyncha. In: Hofrichter, R. (ed.), Das Mittelmeer. Fauna, Flora, Ökologie. Vol. II/2 Bestimmungsführer, pp. 646-653. Spektrum Akademischer Verlag.
note: general treatment of Kinorhyncha and identification guide for Mediterranean species [details] Available for editors  PDF available [request]

biology source Nyholm, K.-G.; Nyholm, P.-G. (1983). 11. Kinorhyncha. In: Adiyodi, K.G. & Adiyodi, R.G. (eds), Reproductive biology of invertebrates, Vol. II: Spermatogenesis and sperm function. John Wiley & Sons, New York. pp. 207-220.
note: The book chapter reviews aspects of the reproductive biology of Kinorhyncha. [details] Available for editors  PDF available [request]

biology source Higgins, R. P. (1974). Chapter 11 Kinorhyncha. In: Giese, A. C. & Pearse, J. S. (Eds), Reproduction of marine invertebrates, Vol. I. Acoelomate and pseudocoelomate metazoans. Academic Press, New York, pp. 507-518.
note: The book chapter reviews the reproduction of Kinorhyncha. [details] Available for editors  PDF available [request]

biology source Neuhaus, B.; Higgins, R. P. (2002). Ultrastructure, biology, and phylogenetic relationships of Kinorhyncha. <em>Integrative and Comparative Biology.</em> 42(3): 619-632., available online at https://doi.org/10.1093/icb/42.3.619
note: review of TEM studies, functional morphology and relationships to Priapulida and Loricifera; new findings about localization of chitin in the cuticle at TEM level [details] 

biology source Neuhaus, B. (1999). Kinorhyncha. In: Knobil, E. & Neill, J. D. (eds), Encyclopedia of reproduction, Vol 2, pp. 933-937. Academic Press, New York.
note: review of reproduction in Kinorhyncha [details] Available for editors  PDF available [request]

biology source Zelinka, C. (1914). Zwei Ektoparasiten der Echinoderen aus der Klasse der Ciliaten. <em>Verhandlungen der Gesellschaft Deutscher Naturforscher und Ärzte.</em> 85. Versammlung zu Wien, Teil 2, 1. Hälfte, Nr. 13, 680-683.
note: Inadvertantly, the article introduces two protozoan ectoparasites on kinorhynchs, viz, Cothurnia bütschlii and Acineta tuba. These species are actually redescribed in Zelinka (1928). [details] OpenAccess publication

biology source Adrianov, A. V.; Malakhov, V. V. (1999). 7. Kinorhyncha. <em>In: Adiyodi, K.G. & Adiyodi, R.G. (eds.) Reproductive biology of invertebrates, Vol. IX, Part A: Progress in male gamete ultrastructure and phylogeny. John Wiley & Sons, Chichester.</em> pp. 193–211.
note: The bok chapter reviews the ultrastructure of the male reproductive system of Kinorhyncha. [details] Available for editors  PDF available [request]
 
 Present  Inaccurate  Introduced: alien  Containing type locality 
From editor or global species database
Classification As stated by Neuhaus (2013, p. 273), Kinorhyncha was also named Echinodera, Echinoderen (in German) and Echinoderida.
"Echinodera" was used by: Zelinka (1928), Blake (1930), Chitwood (1951), and Bacescu (1968).
The German "Echinoderen" was used by: Greeff (1869), Metschnikoff (1869), Pagenstecher (1875), Bütschli (1876), Reinhard (1887), and Zelinka (1896, 1907, 1908, 1912, 1913, 1914).
The Italian "Echinoderi" was used by Panceri (1878).
Both "Echinodera" + "Kinorhyncha" were mentioned by: Remane (1929, 1936), Pearse (1936), and Neuhaus (2013).
"die Echinoderiden" was used by: Zelinka (1894),Schepotieff (1907a-b, 1908).
"Echinoderida" was used by: Abe (1930), Johnston (1938), Nyholm (1947a-c), Lang (1949, 1953), Krishnaswamy (1962, lapsus calami: Echinodereda), and McIntyre (1962).
"Echinoderida" + "Kinorhyncha" were mentioned by Southern (1914) and Moore (1973)
"Echinocephala" = "Igelwürmer" were created by Haeckel (1896).
The remaining sources used "Kinorhyncha" only, introduced by Reinhard (1885), some as early as Hartog (1896). [details]

Classification Neuhaus (2013, p. 273): "Some confusion appears in the literature (Remane 1928, 1929, 1936; Zelinka 1928; Higgins 1971; Adrianov & Malakhov 1999a) concerning the names of and within the taxon Kinorhyncha and the time these names have been introduced (corrected in Tabs 6, 7, 9). Whereas Dujardin (1851) calls his new species simply “l’Echinodère”, Claparède (1863) formally assigns it to a genus and names it Echinoderes dujardinii. No unifying scientific name is provided for species of Echinoderes, probably also because it takes some time to realize that the taxon represents a group of its own. The name “Kinorhyncha (Echinoderes)” is first used in the seldom-cited contribution of Reinhard (1885) in Russian. Subsequently, this very same article is published again in German, almost without any major changes (Reinhard 1887), and this time it receives much more attention, especially in the German-speaking scientific community (Remane 1928, 1929, 1936; Zelinka 1928). Reinhard (1885: p. 296, 1887: p. 463) chooses the name “Kinorhyncha” (kinein = Greek for move, rhynchos = Greek for snout), because he wants to avoid confusion with, e.g., Echinodermata, and because he misidentifies the posterior extensions of the esophagus bulging between the pharynx protractors as brain and consequently the protrusible introvert as protrusible snout of the alimentary canal (Zelinka 1928: p. 38). However, Zelinka (1928: p. 224) latinizes the German name “die Echinoderen” and refers to the Kinorhyncha as “Echinodera” (echinos = Greek for hedgehog, deres = Greek for neck; terms refer to spiny scalids of the introvert, which Dujardin in 1851 obviously interprets as neck, Echinoderes represents the first described genus of Kinorhyncha). Other authors prefer the name “Echinoderida” derived from the first family erected within Kinorhyncha, Echinoderidae (e.g., Johnston 1938; Nyholm 1947c; Lang 1949, 1953; McIntyre 1962). In summary, the first introduced scientific group name is Kinorhyncha, but the morphologically correct name would be Echinodera." [details]

Etymology "Reinhard (1885: p. 296, 1887: p. 463) chooses the name “Kinorhyncha” (kinein = Greek for move, rhynchos = Greek for snout), because he wants to avoid confusion with, e.g., Echinodermata, and because he misidentifies the posterior extensions of the esophagus bulging between the pharynx protractors as brain and consequently the protrusible introvert as protrusible snout of the alimentary canal (Zelinka 1928: p. 38)." [details]

Museum Type and non-type specimens of Kinorhyncha are mainly deposited in the following collections:
1) Museum of Natural History, Smithsonian Institution, Washington D.C., USA (USNM; search colection: https://collections.nmnh.si.edu/search/iz/). The museum houses also a Microsoft Access database of the stations of its former curator R. P. Higgins, which is complementary to the museum's catalogued and uncatalogued kinorhynch specimens. Field numbers by Higgins start with RH-...
2) Zoological Museum, now The Natural History Museum of Denmark, Copenhagen, Denmark (ZMUC-KIN, NHMD; search collection: http://www.daim.snm.ku.dk/The-digitized-type-collection)
3) Zoologisches Museum Berlin, now Museum für Naturkunde Berlin, Germany (ZMB, MNB; search collection: http://zmb.sesam.senckenberg.de/page/index.htm). [details]

Reproduction Higgins (1974) reviewed sexual dimorphism, anatomy of the reproductive system, reproductive behaviour, and postembryonic development of Kinorhyncha and provided some new own data. [details]

Synonymy "Zelinka (1928: p. 224) latinizes the German name “die Echinoderen” and refers to the Kinorhyncha as “Echinodera” (echinos = Greek for hedgehog, deres = Greek for neck; terms refer to spiny scalids of the introvert, which Dujardin in 1851 obviously interprets as neck, Echinoderes represents the first described genus of Kinorhyncha). Other authors prefer the name “Echinoderida” derived from the first family erected within Kinorhyncha, Echinoderidae (e.g., Johnston 1938; Nyholm 1947c; Lang 1949, 1953; McIntyre 1962). In summary, the first introduced scientific group name is Kinorhyncha, but the morphologically correct name would be Echinodera." [details]

Taxonomy Higgins (1986): species list of Kinorhyncha. [details]

Taxonomy Higgins, R. P. (1981): The author described the taxonomic ranks within the phylum Kinorhyncha from order to genus level and provided illustrations of the genera. [details]
    Definitions

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LanguageName 
Dutch stekelwormen  [details]
English spiny-crown wormsmud dragonskinorhynchs  [details]
French Kinorhynques  [details]
German Kinorhynchen, Hakenrüssler, Igelnackentiere, Stoßrüssler, Igelwürmer  [details]
Italian Chinorhinchi  [details]
Japanese 動吻動物門動吻動物  [details]
Romanian Kinorhinchii  [details]
Spanish Cinorrincos, Kinorrincos, Quinorrinchos, Quinorrincos  [details]
Swedish pansarmaskar  [details]
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