Nordheim, Henning von (1989). Six new species of Protodrilus (Annelida, Polychaeta) from Europe and New Zealand, with a concise presentation of the genus. Zoologica Scripta 18 (2), 245–268.
World Polychaeta Database (WPolyDb). Author alphabetises himself as Nordheim, rather than von Nordheim.
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Protodrilus haurakiensis sp.n., P. jouinae sp.n., P. jägersteni sp.n., P. submersus sp.n. and P. litoralis sp.n. from New Zealand and P. gracilis sp.n. from Europe are described. Protodrilus jägersteni, P. submersus and P. litoralis are sibling species. Sperm dimorphism in all species investigated by the author is recorded for the first time in polychaetes. Two of the three so-called 'ecological forms' of P. hypoleucus and P. adhaerens are classified as variations of the type species without a particular taxonomic rank. In a table the partly actualized data of all 30 known Protodrilus species are presented, 8 of which are very insufficiently described. Synonyms in Protodrilus and similarities between P. rubropharyngeus Jägersten and P. flavocapitatus Uljanin are mentioned.