The main theme of my research is the systematics and comparative anatomy of free-living and parasitic copepods, and the application of phylogenetic reconstruction to examine their evolution and ecological radiation, using morphology and molecular markers. Copepods are one of the best models to study fundamental phenomena like the evolution of parasitism and the marine-freshwater transition, and to test fundamental hypotheses such as the claim of oligomerization being the predominant mode of character transformation in Crustacea, and the enemy release hypothesis in invasion ecology. Part of my research focuses on these intrinsically interesting questions.
Teaching students and professionals the methods how to observe, illustrate and describe copepods defines the red line throughout my career. Training the next generation is key to the survival and success of our research community and I have devoted much time to it which brought me great satisfaction. I was involved as a tutor or organizer in student training workshops on free-living and symbiotic copepods since 2001. In 2022 I co-organized the international copepod conference (ICOC) which was widely considered as the most successful ICOC ever, welcoming an unprecedentedly high participation of students. I have served the World Association of Copepodologists (WAC) as a member of the Executive Council (1996–2002; 2022–present, and as Vice-President (2008–2011), President (2011–2014) and Past-President (2014–2017).
In 2017 I became the recipient of the Monoculus Award for research excellence and exceptionally devoted service and contributions to the activities of WAC.
For the last ten years I have assisted the editors of WoRMS in solving nomenclatural issues related to copepods and related crustaceans.
Boxshall, G.A.; Huys, R. (2024). World List of Tantulocarida. Accessed at https://marinespecies.org/aphia.php?p=taxdetails&id=1083 on yyyy-mm-dd, more
Huys, R. & De Smet, G. (1993). Copepoda from the Dutch Continental Shelf, spring 1993. Ghent University, Department of Biology, Marine Biology Section (MARBIOL), Belgium., more
Huys, R. (1984). Copepods from a sublittoral sandy station in the North Sea. Ghent University, Department of Biology, Marine Biology Section (MARBIOL), Belgium., more
Back, J.; Huys, R.; Lee, W. (2010). A new species of the genus Tegastes (Copepoda: Harpacticoida: Tegastidae) from hydrothermal vents in the Okinawa Trough. Zool. Sci. 27(8): 678-688. http://dx.doi.org/10.2108/zsj.27.678
El Nagar, A.; Huys, R.; Bishop, J.D.D. (2010). Widespread occurrence of the Southern Hemisphere ascidian Corella eumyota Traustedt, 1882 on the Atlantic coast of Iberia. Aquat. Invasions 5(2): 169-173. http://dx.doi.org/10.3391/ai.2010.5.2.06
Huys, R.; Kihara, T.C. (2010). Systematics and phylogeny of Cristacoxidae (Copepoda, Harpacticoida): a review. Zootaxa 2568(2568): 1-38
Selden, P.A.; Huys, R.; Stephenson, M.H.; Heward, A.P.; Taylor, P.N. (2010). Crustaceans from bitumen clast in Carboniferous glacial diamictite extend fossil record of copepods. Nature Comm. 1(50): 6 pp. http://dx.doi.org/10.1038/ncomms1049
Huys, R. (2009). Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa, 2183(2183). Magnolia Press: Auckland. ISBN 978-1-86977-400-4. 99 pp.
Huys, R. (2009). On the junior subjective synonyms of Coullia Hamond, 1973 (Copepoda, Harpacticoida, Laophontidae): an update and key to species and related genera. ZooKeys 5(5): 33-40. http://dx.doi.org/10.3897/zookeys.5.64
Huys, R.; Lee, W. (2009). Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae). ZooKeys 23(23): 1-38. http://dx.doi.org/10.3897/zookeys.23.168
Kihara, T.C.; Huys, R. (2009). A new genus of Ectinosomatidae (Copepoda, Harpacticoida) from sublittoral sediments in Ubatuba, São Paulo State, Brazil, including an updated key to genera and notes on Noodtiella Wells, 1965. ZooKeys 17(17): 57-88. https://dx.doi.org/10.3897/zookeys.17.202
Kihara, T.C.; Huys, R. (2009). Corrigenda: Kihara TC & Huys R (2009) A new genus of Ectinosomatidae (Copepoda, Harpacticoida) from sublittoral sediments in Ubatuba, São Paulo State (Brazil), an updated key to genera and notes on Noodtiella Wells, 1965. ZooKeys 17: 57-88. ZooKeys 18(Special Issue): 181-182. dx.doi.org/10.3897/zookeys.18.249
Kihara, T.C.; Huys, R. (2009). Contributions to the taxonomy of the Normanellidae (Copepoda, Harpacticoida): description of a new genus from the Brazilian continental shelf and re-assignment of Pseudocletodes vararensis Scott & Scott, 1893 (ex Nannopodidae). Zootaxa 2233(2233): 1-38
Huys, R.; Mu, F.-H. (2008). Description of a new species of Onychostenhelia Itô (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa 1706: 51-68
Sak, S.; Karaytug, S.; Huys, R. (2008). A review of Pseudoleptomesochrella Lang, 1965 (Copepoda, Harpacticoida, Ameiridae), including a redescription of P. halophila (Noodt, 1952) from the Black Sea and a key to species. Zootaxa 1758: 45-60
Sak, S.; Karaytug, S.; Huys, R. (2008). Ciplakastacus gen. nov., a primitive genus of Leptastacidae (Copepoda, Harpacticoida) from the Mediterranean coast of Turkey. J. Nat. Hist. 42(37-38): 2443-2459. http://dx.doi.org/10.1080/00222930802277632
Huys, R.; LLewellyn-Hughes, J.; Conroy-Dalton, S.; Olson, P.D.; Spinks, J.N.; Johnston, D.A. (2007). Extraordinary host switching in siphonostomatoid copepods and the demise of the Monstrilloida: Integrating molecular data, ontogeny and antennulary morphology. Mol. Phylogenet. Evol. 43(2): 368-378. http://dx.doi.org/10.1016/j.ympev.2007.02.004
Huys, R.; LLewellyn-Hughes, J.; Olson, P.D.; Nagasawa, K. (2006). Small subunit rDNA and Bayesian inference reveal Pectenophilus ornatus (Copepoda incertae sedis) as highly transformed Mytilicolidae, and support assignment of Chondracanthidae and Xarifiidae to Lichomolgoidea (Cyclopoida). Biol. J. Linn. Soc. 87(3): 403-425. http://dx.doi.org/10.1111/j.1095-8312.2005.00579.x
Huys, R.; Conroy-Dalton, S. (2006). Revision of the genus Evansula T. Scott, 1906 (Copepoda, Harpacticoida, Cylindropsyllidae) with a description of three new species. Zool. J. Linn. Soc. 147(4): 419-472. http://dx.doi.org/10.1111/j.1096-3642.2006.00227.x
Huys, R.; Conroy-Dalton, S. (2006). Comparative analysis of vermiform genera (Copepoda: Harpacticoida: Cylindropsyllidae), including descriptions of Boreovermis, new genus and Bolbotelos, new genus, from the Southern North Sea. J. Crust. Biol. 26(2): 206-223. http://dx.doi.org/10.1651/C-2609.1
Huys, R.; Ohtsuka, S.; Conroy-Dalton, S.; Kikuchi, Y. (2005). Description of two new species of Neotachidius Shen & Tai, 1963 (Copepoda, Harpacticoida, Tachidiidae) from Korean brackish waters and proposal of a new genus for Tachidius (Tachidius) vicinospinalis Shen & Tai, 1964. Zool. J. Linn. Soc. 143(1): 133-159. http://dx.doi.org/10.1111/j.1096-3642.2004.00148.x
Ivanenko, V.N.; Defaye, D.; Huys, R. (2005). A new species of Ambilimbus nom. nov., a replacement name for Amphicrossus Huys, 1991 (Copepoda, Cyclopoida, Erebonasteridae), from the Mid-Atlantic Ridge at 36°N (hydrothermal vent site Rainbow). Mar. Biol. Res. 1(3): 233-241. http://dx.doi.org/10.1080/17451000510019097
Böttger-Schnack, R.; Huys, R. (2004). Size polymorphism in Oncaea venusta Philippi, 1843 and the validity of O. frosti Heron, 2002: a commentary. Hydrobiologia 513: 1-5. http://dx.doi.org/10.1023/B:hydr.0000018301.50664.3d
Huys, R.; Joon Song, S. (2004). The Ismardiidae Leigh-Sharpe (Copepoda, Harpacticoida incertae sedis): Enigmatic as ever? J. Crust. Biol. 24(1): 37-53. http://dx.doi.org/10.1651/C-2412
Karaytug, S.; Huys, R. (2004). Taxonomic position of and generic distinction between Parepactophanes Kunz, 1935 and Taurocletodes Kunz, 1975 (Copepoda, Canthocamptidae incertae sedis), with description of a new species from the Black Sea. Zool. J. Linn. Soc. 140(4): 469-486. http://dx.doi.org/10.1111/j.1096-3642.2003.00101.x
Huys, R. (2003). Book review: Martin, Joel W., and George E. Davis. 2001. An Updated Classification of the Recent Crustacea. Natural History Museum of Los Angeles County Contributions in Science 39: 1-124. J. Crust. Biol. 23(2): 495-497. http://dx.doi.org/10.1651/0278-0372(2003)023[0495:BR]2.0.CO;2
Huys, R.; López-Gonzáles, P.J.; Roldán, E.; Luque, A. (2002). Brooding in cocculiniform limpets (Gastropoda) and familial distinctiveness of the Nucellicolidae (Copepoda): misconceptions reviewed from a chitonophilid perspective. Biol. J. Linn. Soc. 75(2): 187-217. http://dx.doi.org/10.1046/j.1095-8312.2002.00010.x
Mu, F.-H.; Huys, R. (2002). New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cah. Biol. Mar. 43(2): 179-206
Böttger-Schnack, R.; Huys, R. (2001). Taxonomy of Oncaeidae (Copepoda, Poecilostomatoida) from the Red Sea: III. Morphology and phylogenetic position of Oncaea subtilis Giesbrecht, 1892. Hydrobiologia 453-454: 467-481. http://dx.doi.org/10.1007/0-306-47537-5_40
Huys, R. (2001). Splanchnotrophid systematics: a case of polyphyly and taxonomic myopia. J. Crust. Biol. 21(1): 106-156
Huys, R.; Boxshall, G. (2001). An appreciation of the contribution of Arthur Humes to copepod systematics. J. Crust. Biol. 21(1): 13-27. http://dx.doi.org/10.1163/20021975-99990106
Lee, W.; Huys, R. (2001). Erratum. New Aegisthidae (Copepoda: Harpacticoida) from western Pacific cold seeps and hydrothermal vents. Zool. J. Linn. Soc. 131: 249
Ohtsuka, S.; Huys, R. (2001). Sexual dimorphism in calanoid copepods: morphology and function. Hydrobiologia 453-454: 441-466. http://dx.doi.org/10.1023/A:1013162605809
Lee, W.; Huys, R. (2000). New Aegisthidae (Copepoda: Harpacticoida) from western Pacific cold seeps and hydrothermal vents. Zool. J. Linn. Soc. 129: 1-171. http://dx.doi.org/10.1006/zjls.1999.0197, more
López-González, P.J.; Bresciani, J.; Huys, R.; González, Á.F.; Guerra, Á.; Pascual, S. (2000). Description of Genesis vulcanoctopusi gen. et sp. nov. (Copepoda: Tisbidae) parasitic on a hydrothermal vent octopod and a reinterpretation of the life cycle of cholidyinid harpacticoids. Cah. Biol. Mar. 41(3): 241-253
Conroy-Dalton, S.; Huys, R. (1999). New genus of Aegisthidae (Copepoda: Harpacticoida) from hydrothermal vents on the Galapagos Rift. J. Crust. Biol. 19(2): 408-431
Huys, R.; Lee, W. (1999). On the relationships of the Normanellidae and the recognition of Cletopsyllidae grad. nov. (Copepoda, Harpacticoida). Zool. Anz. 237: 267-290
Lee, W.; Huys, R. (1999). New Tachidiella (Copepoda, Harpacticoida, Tisbidae) from the Antarctic and Norway including a review of the genus. Zoosystema 21(3): 419-444
Lee, W.; Huys, R. (1999). New Normanellidae (Copepoda: Harpacticoida) from western Pacific cold seeps including a review of the genus Normanella. Cah. Biol. Mar. 40(3): 203-262
Lee, W.; Huys, R. (1999). Bathylaophonte gen. nov. from deep-sea hydrothermal vents and the polyphyly of Paronychocamptus (Copepoda : Harpacticoida). Cah. Biol. Mar. 40(4): 293-328
Böttger-Schnack, R.; Huys, R. (1998). Species groups within the genus Oncaea Philippi, 1843 (Copepoda, Poecilostomatoida). J. Mar. Syst. 15(1-4): 369-371. http://dx.doi.org/10.1016/S0924-7963(97)00072-9
Boxshall, G.A.; Huys, R. (1998). The ontogeny and phylogeny of copepod antennules. Phil. Trans. R. Soc. Lond. (B Biol. Sci.) 353(1369): 765-786. http://dx.doi.org/10.1098/rstb.1998.0242
Conroy-Dalton, S.; Huys, R. (1998). Towards a revision of Ameira Boeck, 1865 (Harpacticoida, Ameiridae): reinstatement of Psammameira Noodt, 1952. Zoologica Scri. 27(3): 247-261. http://dx.doi.org/10.1111/j.1463-6409.1998.tb00439.x
Böttger-Schnack, R.; Huys, R. (1997). Archioncaea arabica gen. et sp. nov., a remarkable oncaeid (Copepoda: Poecilostomatoida) from the northern Arabian Sea. Cah. Biol. Mar. 38(2): 79-89
Huys, R.; Møbjerg, N.; Kristensen, R.M. (1997). New tantulocarid Polynyapodella ambrosei gen. et sp., (Basipodellidae) from the Northeast Water Polynya (Greenland) with emphasis on the phylogeny of its host genus Cervinia (Copepoda: Harpacticoida). Cah. Biol. Mar. 38(3): 181-199
Huys, R.; Böttger-Schnack, R. (1997). On the diphyletic origin of the Oncaeidae Giesbrecht, 1892 (Copepoda: Poecilostomatoida) with a phylogenetic analysis of the Lubbockiidae fam. nov. Zool. Anz. 235: 243-261
Huys, R.; Conroy-Dalton, S. (1997). Discovery of hydrothermal vent Tantulocarida on a new genus of Argestidae (Copepoda: Harpacticoida). Cah. Biol. Mar. 38(4): 235-249
Huys, R.; Todaro, M.A. (1997). Meloriastacus ctenidis gen. et sp. nov: A primitive interstitial copepod (Harpacticoida, Leptastacidae) from Tuscany. Ital. J. Zoolog. 64(2): 181-196
Conroy-Dalton, S.; Huys, R. (1996). Towards a revision of Ameira Boeck, 1865 (Harpacticoida, Ameiridae): re-examination of the A. tenella-group and the establishment of Filexilia gen. n. and Glabrameira gen. n. Zoologica Scri. 25(4): 317-339. http://dx.doi.org/10.1111/j.1463-6409.1996.tb00169.x
Huys, R. (1996). Superornatiremidae fam. nov. (Copepoda: Harpacticoida): An enigmatic family from North Atlantic anchihaline caves. Sci. Mar. (Barc.) 60(4): 497-542
Huys, R. (1996). Biuncus nom. nov., a replacement name for Singularia Huys, 1995 (Copepoda: Harpacticoida: Paramesochridae). J. Nat. Hist. 30: 1261. http://dx.doi.org/10.1080/00222939600770691
Huys, R.; Bodiou, J.-Y.; Bodin, P. (1996). A revision of Psamathea (Harpacticoida: Leptastacidae) with description of P. brittanica spec. nov. Vie Milieu (1980) 46(1): 7-19
Huys, R.; Conroy-Dalton, S. (1996). A revision of Leptopontia T. Scott (Copepoda: Harpacticoida) with description of six new species. Zool. J. Linn. Soc. 118(3): 197-239. http://dx.doi.org/10.1111/j.1096-3642.1996.tb01265.x, more
Huys, R.; Gee, J.M. (1996). Sentiropsis, Peltisenia and Afrosenia: three new genera of Paranannopidae (Copepoda, Harpacticoida). Cah. Biol. Mar. 37(1): 49-75
Huys, R.; Gee, J.M. (1996). Prionos gen. nov. from the meiofauna of a Malaysian mangrove forest and the status of Psammis borealis (Copepoda, Harpacticoida, Paranannopidae). Cah. Biol. Mar. 37(3): 227-248
Huys, R. (1995). Some remarks on the taxonomic status of Paraschizopera Wells, 1981 (Copepoda: Harpacticoida). Hydrobiologia 308(1): 23-28. http://dx.doi.org/10.1007/BF00037783
Huys, R. (1995). A new genus of Paramesochridae (Copepoda: Harpacticoida) from amphioxus-sand, Elat, Israel. J. Nat. Hist. 29(3): 673-684. http://dx.doi.org/10.1080/00222939500770231
Boxshall, G.; Huys, R. (1994). Asterocheres reginae, a new species of parasitic copepod (Siphonostomatoida: Asterocheridae) from a sponge in Belize. Syst. Parasitol. 27(1): 19-33. http://dx.doi.org/10.1007/BF02185665
Gee, J.M.; Huys, R. (1994). Paranannopidae (Copepoda: Harpacticoida) from sublittoral soft sediments in Spitsbergen. J. Nat. Hist. 28: 1007-1046
Huys, R. (1993). Styracothoracidae (Copepoda: Harpacticoida), a new family from the Philippine deep sea. J. Crust. Biol. 13(4): 769-783, more
Huys, R.; Boxshall, G.; Casnova, J.-P. (1993). A redescription of Cumoniscus Bonnier, 1903 (Crustacea: Tantulocarida) from a cumacean host in the Bay of Biscay. Syst. Parasitol. 26(1): 75-79. http://dx.doi.org/10.1007/BF00009651
Huys, R.; Boxshall, G.A.; Lincoln, R.J. (1993). The tantulocaridan life cycle: the circle closed? J. Crust. Biol. 13(3): 432-442. https://dx.doi.org/10.2307/1548786
Huys, R.; Willems, K. (1993). A revision of Cylinula and two new species of Cylindropsyllus (Copepoda, Harpacticoida, Cylindropsyllidae). Zoologica Scri. 22(4): 347-362. http://dx.doi.org/10.1111/j.1463-6409.1993.tb00364.x
Boxshall, G.; Huys, R. (1992). A homage to homology: patterns of copepod evolution. Acta Zool. (Stockh.) 73(5): 327-334
Humes, A.; Huys, R. (1992). Copepoda (Poecilostomatoida and Siphonostomatoida) from deep-sea hydrothermal vent areas off British Columbia, including Amphicrossus altalis, a new species of Erebonasteridae, with notes on the taxonomic position of the genus Tychidion Humes. Can. J. Zool. 70: 1369-1380. https://dx.doi.org/10.1139/z92-193
Huys, R.; Herman, P.M.J.; Heip, C.H.R.; Soetaert, K. (1992). The meiobenthos of the North Sea: density, biomass trends and distribution of copepod communities. ICES J. Mar. Sci./J. Cons. int. Explor. Mer 49(1): 23-44, more
Huys, R.; Ohtsuka, S.; Boxshall, G.A.; Ito, T. (1992). Itoitantulus misophricola gen. et sp. nov.: First record of Tantulocarida (Crustacea: Maxillopoda) in the North Pacific Region. Zool. Sci. 9: 875-886
Huys, R.; Gee, J.M. (1992). A revision of Danielssenia perezi Monard, D. paraperezi Soyer, D. eastwardae Coul (Harpacticoida: Paranannopidae) and their transfer to a new genus. Zool. J. Linn. Soc. 104: 31-56
Ohtsuka, S.; Huys, R.; Boxshall, G.A.; Ito, T. (1992). Misophriopsis okinawensis sp. nov. (Crustacea: Copepoda) from hyperbenthic waters off Okinawa, south Japan, with definitions of related genera Misophria Boeck, 1864 and Stygomisophria gen. nov. Zool. Sci. 9: 859-874
Gee, J.M.; Huys, R. (1991). A review of Paranannopidae (Copepoda: Harpacticoida) with claviform aesthetascs on oral appendages. J. Nat. Hist. 25(5): 1135-1169. https://dx.doi.org/10.1080/00222939100770741, more
Huys, R. (1991). Tantulocarida (Crustacea: Maxillopoda): a new taxon from the temporary meiobenthos. Mar. Ecol. (Berl.) 12(1): 1-34
Smol, N.; Huys, R.; Vincx, M. (1991). A 4-years' analysis of the meiofauna community of a dumping site for TiO2-waste off the Dutch coast. Chem. Ecol. 5: 197-215. http://dx.doi.org/10.1080/02757549108035250
Gee, J.M.; Huys, R. (1990). The rediscovery of Danielssenia intermedia Wells 1965 (Copepoda, Harpacticoida): a missing link between the "danielsseniid" genera and Paranannopus Long 1936 (Paranannopidae). J. Nat. Hist. 24(6): 1549-1571. https://dx.doi.org/10.1080/00222939000770891, more
Huys, R. (1990). A new harpacticoid copepod family collected from Australian sponges and the status of the subfamily Rhynchothalestrinae Lang. Zool. J. Linn. Soc. 99(1): 51-115. http://dx.doi.org/10.1111/j.1096-3642.1990.tb01222.x, more
Huys, R. (1990). Pholenota spatulifera Vervoort (Copepoda: Harpacticoida): aberrant laophontid or specialized diosaccid? J. Nat. Hist. 24: 653-646. https://dx.doi.org/10.1080/00222939000770431
Huys, R. (1990). Campyloxiphos dineti gen. et spec. nov. from off Namibia and a redefinition of the Deoterthridae Boxshall and Lincoln (Crustacea: Tantulocarida). J. Nat. Hist. 24(2): 415-432. https://dx.doi.org/10.1080/00222939000770311
Huys, R. (1990). Adenopleurella, new genus, Proceropes, new genus, Sarsocletodes Wilson (ex Laophontidae), and Miroslavia Apostolov (ex Cletodidae): reprensentatives of a new family (Copepoda: Harpacticoida). J. Crust. Biol. 10(2): 340-363, more
Huys, R.; Boxshall, G. (1990). Discovery of Centobnaster humesi, new genus, new species (Erebonasteridae), the most primitive poecilostomatoid copepod known, in New Caledonian deep waters. J. Crust. Biol. 10(3): 504-519. https://dx.doi.org/10.2307/1548341
Boxshall, G.; Huys, R.; Lincoln, R. (1989). A new species of Microdajus (Crustacea: Tantulocarida) parasitic on a tanaid in the north-eastern Atlantic, with observations on M. langi Greve. Syst. Parasitol. 14(1): 17-30. http://dx.doi.org/10.1007/BF00019991
Boxshall, G.; Huys, R. (1989). New tantulocarid, Stygotantulus stocki, parasitic on harpacticoid copepods, with an analysis of the phylogenetic relationships within the Maxillopoda. J. Crust. Biol. 9(1): 126-140. https://dx.doi.org/10.2307/1548454
Huys, R.; Coomans, A. (1989). Echinoderes higginsi sp. n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparède. Zoologica Scri. 18(2): 211-221. http://dx.doi.org/10.1111/j.1463-6409.1989.tb00446.x
Huys, R.; Thistle, D. (1989). Bathycamptus eckmani gen. et spec. nov. (Copepoda, Harpacticoida) with a review of the taxonomic status of certain other deepwater harpacticoids. Hydrobiologia 185(2): 101-126. http://dx.doi.org/10.1007/BF00010809, more
Heip, C.H.R.; Warwick, R.M.; Carr, M.R.; Herman, P.M.J.; Huys, R.; Smol, N.; Van Holsbeke, K. (1988). Analysis of community attributes of the benthic meiofauna of Frierfjord-Langesundfjord. Mar. Ecol. Prog. Ser. 46(1-3): 171-180. http://dx.doi.org/10.3354/meps046171
Huys, R. (1988). A redescription of the presumed associated Caligopsyllus primus Kunz, 1975 (Harpacticoida, Paramesochridae) with emphasis on its phylogenetic affinity with Apodopsyllus Kunz, 1962. Hydrobiologia 162(1): 3-19. http://dx.doi.org/10.1007/BF00014330
Huys, R. (1988). On the identity of the Namakosiramiidae Ho & Perkins 1977 (Crustacea, Copepoda), including a review of harpacticoid associates of Echinodermata. J. Nat. Hist. 22(6): 1517-1532. https://dx.doi.org/10.1080/00222938800770941
Huys, R. (1988). Studies on the Cylindropsyllidae (Copepoda, Harpacticoida): 2. A revision of the genus Cylindropsyllus Brady. Zoologica Scri. 17(3): 253-276. http://dx.doi.org/10.1111/j.1463-6409.1988.tb00100.x
Huys, R. (1987). Studies on the Cylindropsyllidae (Copepoda, Harpacticoida): 1. The status of Leptastacus laticaudatus Nicholls. Zoologica Scri. 16(2): 155-166. http://dx.doi.org/10.1111/j.1463-6409.1987.tb00062.x
Huys, R. (1987). Paramesochra T. Scott, 1892 (Copepoda, Harpacticoida): a revised key, including a new species from the SW Dutch coast and some remarks on the phylogeny of the Paramesochridae. Hydrobiologia 144(3): 193-210. http://dx.doi.org/10.1007/BF00005554, more
Huys, R.; Clark, P.F. (2009). Case 3467. Zosimidae Seifried, 2003 (Crustacea, Copepoda, Harpacticoida): proposed emendation of spelling to Zosimeidae to remove homonymy with Zosiminae Alcock, 1898 (Crustacea, Decapoda, Xanthidae). Bull. Zool. Nomencl. 66(1): 24-29
Willems, W.R.; Curini-Galletti, M.; Ferrero, T.J.; Fontaneto, D.; Heiner, I.; Huys, R.; Ivanenko, V.N.; Kristensen, R.M.; Kånneby, T.; MacNaughton, M.O.; Martinéz Arbizu, P.; Todaro, M.A.; Sterrer, W.; Jondelius, U. (2009). Meiofauna of the Koster-area, results from a workshop at the Sven Lovén Centre for Marine Sciences (Tjärnö, Sweden). Meiofauna Mar. 17: 1-34
Huys, R. (2008). Case 3470. Halectinosoma Vervoort, 1962 (Crustacea, Copepoda, Harpacticoida): proposed conservation of usage by designation of Ectinosoma chrystalii Scott, 1894 as the type species and by giving it precedence over Pararenosetella Lang, 1944. Bull. Zool. Nomencl. 65(4): 276-281
Huys, R. (2008). Case 3471. Heterolaophonte Lang, 1948 (Crustacea, Copepoda, Harpacticoida): proposed precedence over Mesolaophonte Nicholls, 1941 and Monolaophonte Nicholls, 1941. Bull. Zool. Nomencl. 65(4): 282-287
Lee, W.; Huys, R. (2002). A new genus of groundwater Ameiridae (Copepoda, Harpacticoida) from boreholes in Western Australia and the artificial status of Stygonitocrella Petkovski, 1976. Bull. Nat. Hist. Mus., Zool. Ser. 68(1): 39-50
Huys, R.; Conroy-Dalton, S. (2000). Generic concepts in the Clytemnestridae (Copepoda, Harpacticoida), revision and revival. Bull. Nat. Hist. Mus., Zool. Ser. 66(1): 1-48
Huys, R.; Lee, W. (2000). Basal resolution of laophontid phylogeny and the paraphyly of Esola Edwards. Bull. Nat. Hist. Mus., Zool. Ser. 66(1): 49-107
Bouck, L.; Thistle, D.; Huys, R. (1999). Systematics and phylogeny of Zausodes C.B. Wilson, 1932 (Copepoda, Harpacticoida, Harpacticidae), including three new species from the northern Gulf of Mexico. Bull. Nat. Hist. Mus., Zool. Ser. 65(2): 73-122
Todaro, M.A.; Huys, R. (1998). La meiofauna delle Secche della Meloria: osservazioni su un nuovo Leptastacidae (Copepoda, Harpacticoida) = Meiofauna from the Meloria Shoals: observations on a new Leptastacidae (Copepoda, Harpacticoida). Biol. Mar. Medit. 5(1): 591-595
Böttger-Schnack, R.; Huys, R. (1997). Morphological observations on Oncaea mediterranea (Claus, 1863) (Copepoda, Poecilostomatoida) with a comparison of Red Sea and eastern Mediterranean populations. Bull. Nat. Hist. Mus., Zool. Ser. 63(2): 137-147
Gee, J.M.; Huys, R. (1996). An appraisal of the taxonomic position of Enhydrosoma buchholzi (Boeck, 1872), E. bifurcarostratum Shen & Tai, 1965, E.barnishi Wells, 1967 and E. vervoorti Fiers, 1987 with definition of two new genera (Copepoda, Harpacticoida, Cletodidae). Sarsia 81: 161-191, more
Huys, R.; Ohtsuka, S.; Boxshall, G.A. (1994). A new tantulocaridan (Crustacea: Maxillopoda) parasitic on calanoid, harpacticoid and cyclopoid copepods. Publ. Seto Mar. Biol. Lab. 36(4): 197-209. hdl.handle.net/2433/176239
Huys, R.; Conroy-Dalton, S. (1993). Willemsia gen. nov. and Boreopontia Willems, 1981 revisited (Harpacticoida: Cylindropsyllidae). Sarsia 78(3-4): 273-300. http://dx.doi.org/10.1080/00364827.1993.10413541, more
Huys, R.; Gee, M. (1993). A revision of Danielssenia Boeck and Psammis Sars with the establishment of two new genera Archisenia and Bathypsammis (Harpacticoida: Paranannopidae). Bull. Nat. Hist. Mus., Zool. Ser. 59(1): 45-81, more
Huys, R.; Ohtsuka, S. (1993). A new species of Syrticola Willems & Claeys, 1982 (Copepoda: Harpacticoida) from Japan with notes on the type species. Bull. Nat. Hist. Mus., Zool. Ser. 59(1): 83-94
Heip, C.H.R.; Basford, D.; Craeymeersch, J.A.; Dewarumez, J.-M.; de Wilde, P.; Dörjes, J.; Duineveld, G.; Eleftheriou, A.; Herman, P.M.J.; Huys, R.; Irion, G.; Niermann, U.; Kingston, P.; Künitzer, A.; Rachor, E.; Rumohr, H.; Soetaert, K.; Soltwedel, T. (1992). The benthic communities of the North Sea: a summary of the results of the North Sea benthos survey. ICES Coop. Res. Rep. 190: 148-175, more
Heip, C.H.R.; Huys, R.; Alkemade, R. (1992). Community structure and functional roles of meiofauna in the North Sea. Neth. J. Aquat. Ecol. 26(1): 31-41. http://dx.doi.org/10.1007/BF02298026
Huys, R. (1992). The amphiatlantic distribution of Leptastacus macronyx (T. Scott, 1892) (Copepoda: Harpacticoida): a paradigm of taxonomic confusion, and a cladistic approach to the classification of the Leptastacidae Lang, 1948. Meded. Kon. Acad. Wet. Lett. Kunst. Klasse der Wet. 54(4): 21-196, more
Huys, R.; Andersen, P.; Kristensen, R.M. (1992). Tantulacus hoegi gen. et sp. nov. (Tantulocarida: Deoterthridae) from the meiobenthos of the Faroe Bank, North Atlantic. Sarsia 76(4): 287-297. http://dx.doi.org/10.1080/00364827.1992.10413486
Huys, R.; Boxshall, G.A.; Böttger-Schnack, R. (1992). On the discovery of the male of Mormonilla Giesbrecht, 1891 (Copepoda: Mormonilloida). Bull. Br. Mus. Nat. Hist., Zool. 58(2): 157-170
Huys, R. (1991). Crustacea Copepoda: Amphicrossus pacificus gen. et sp. nov., an erebonasterid copepod (Poecilostomatoida) from the New Caledonian continental shelf, in: Crosnier, A. (Ed.) Résultats des Campagnes MUSORSTOM 9. Mémoires du Muséum national d'Histoire naturelle. Série A, Zoologie, 152: pp. 63-77
Huys, R.; Boxshall, G.A. (Ed.) (1991). Copepod evolution. Ray Society (Publications), 159. Ray Society: London. ISBN 0-903-87421-0. 468 pp.
Heip, C.H.R.; Huys, R.; Vincx, M.; Vanreusel, A.; Smol, N.; Herman, R.; Herman, P.M.J. (1990). Composition, distribution, biomass and production of North Sea meiofauna. Neth. J. Sea Res. 26(2-4): 333-342. http://dx.doi.org/10.1016/0077-7579(90)90095-X
Huys, R. (1990). Allocation of the Mantridae Leigh-Sharpe to the Cyclopoida (Crustacea: Copepoda) with notes on Nearchinotodelphys Ummerkutty. Bijdr. Dierkd. 60(3-4): 283-291
Huys, R. (1990). Amsterdam Expeditions to the West Indian Islands, Report 64. A new family of harpacticoid copepods and an analysis of the phylogenetic relationships within the Laophontoidea T. Scott. Bijdr. Dierkd. 60(2): 79-120
Huys, R.; Boxshall, G. (1990). The rediscovery of Cyclopicina longifurcata (Scott) (Copepoda: Cyclopinidae) in deep water in the North Atlantic, with a key to genera of the subfamily Cyclopininae. Sarsia 75(1): 17-32. http://dx.doi.org/10.1080/00364827.1990.10413438
Huys, R. (1989). Dicrotrichura tricincta gen. et spec. nov.: a new tantulocaridan (Crustacea: Maxillopoda) from the Mediterranean deep waters off Corsica. Bijdr. Dierkd. 59(4): 243-249
Huys, R.; Willems, K. (1989). Laophontopsis Sars and the taxonomic concept of the Normanellinae (Copepoda: Harpacticoida): a revision. Bijdr. Dierkd. 59(4): 203-277, more
Huys, R. (1988). Sexual dimorphism in Aegisthid cephalosomic appendages (Copepoda, Harpacticoida): a reappraisal. Bijdr. Dierkd. 58(1): 14-136
Huys, R.; Boxshall, G. (1988). A new genus and species of Tantulocarida (Crustacea: Tantulocarida) parasitic on a harpacticoid copepod from the Skagerrak. Sarsia 73(3): 205-211. http://dx.doi.org/10.1080/00364827.1988.10413407
Huys, R. (1987). Some morphological observations on the Neobradyidae Olofsson, 1917 (Copepoda, Harpacticoida) including the redescription of Antarcticobradya tenuis (Brady, 1910) comb. nov. Bull. K. Belg. Inst. Nat. Wet. 57: 133-148, more
Huys, R.; Herman, R.L.; Heip, C. (1986). Seasonal fluctuations in vertical distribution and breeding activity of a subtidal harpacticoid community in the Southern Bight, North Sea. Neth. J. Sea Res. 20(4): 375-383. http://dx.doi.org/10.1016/0077-7579(86)90004-9
Huys, R.; Gee, J.M.; Moore, C.G.; Hamond, R. (1996). Marine and brackish water harpacticoid copepods part 1: keys and notes for identification of the species. Synopses of the British Fauna, N.S. 51. Field Studies Council: Shrewsbury. ISBN 1-85153-256-0. VII, 352 pp., more
Huys, R. (1991). The meiobenthos of the North Sea. Final report. Delta Institute for Hydrobiological Research: Yerseke.
Huys, R. (2001). Copepoda - Harpacticoida, in: Costello, M.J. et al. (Ed.) European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 268-280, more
Heip, C.H.R.; Hamerlynck, O.; Herman, P.M.J.; Herman, R.; Huys, R.; Vincx, M.; Vranken, G. (1985). Meiobenthos in the Southern Bight of the North Sea, in: Van Grieken, R. et al. (Ed.) Progress in Belgian Oceanographic Research: Proceedings of a symposium held at the Palace of Academies Brussels, 3-5 March 1985. pp. 264-266
Huys, R.; Herman, R. (1985). Seasonal fluctuations in vertical distribution and breeding activity of a subtidal harpacticoid community in the Southern Bight, North Sea, in: Van Grieken, R. et al. (Ed.) Progress in Belgian Oceanographic Research: Proceedings of a symposium held at the Palace of Academies Brussels, 3-5 March 1985. pp. 279-291
Martens, P.M.; Jouk, P.; Huys, R.; Herman, R. (1985). Short note on the relative abundance of the Turbellaria in the meiofauna of sandy habitats in the Southern Bight of the North Sea and on the Belgian beaches, in: Van Grieken, R. et al. (Ed.) Progress in Belgian Oceanographic Research: Proceedings of a symposium held at the Palace of Academies Brussels, 3-5 March 1985. pp. 341-342
Willems, K.; Huys, R. (1988). Harpacticoida (Copepoda, Crustacea) van de Belgisch-Nederlandse kustzone en aangrenzende estuaria: stand van zaken en perspectieven, in: Wouters, K. et al. (Ed.) Symposium Invertebraten van België, samenvattingen, 25-26 nov. 1988. pp. 103, more
Huys, R.; De Smet, G. (1991). Het meiobenthos van het Nederlands Continentaal Plat voorjaar 1991: rapportage in het kader van het Biologisch Monitoring Project. NIOO: Yerseke. [22], 57 pp.
Huys, R.; Heip, C.H.R.; Herman, P.M.J.; Soetaert, K. (1990). The meiobenthos of the North Sea: preliminary results of the North Sea Benthos Survey. CM Documents - ICES, C.M. 1990(Mini:8). ICES: Copenhagen. 17 pp., more
Smol, N.; Huys, R.; Vincx, M. (1989). Studie van het meiobenthos van een dumpingsgebied van titaandioxide-afval in de Nederlands kustwateren, periode 1986-1987. Rijksuniversiteit Gent, Laboratorium voor Morfologie en Systematiek der Dieren, Sectie Mariene Biologie: Gent. 64, 32 tables, 38 graphs pp.
Huys, R.; Mommaerts, J.P.; Steyaert, M.; Van Beveren, C. (1972). Phytoplancton: rapport de synthèse. Synthesis Reports CIPS. KUL/ULB/VUB: Leuven. 8 + tableaux + figures pp.
Huys, R. (1990). Coralliotantulus coomansi gen. et sp.n.: first record of a tantulocaridan (Crustacea: Maxillopoda) from shallow subtidal sands in tropical waters. Stygologia 5(3): 183-198
Boxshall, G.; Huys, R. (1989). New family of deep-sea planktonic copepods, the Paralubbockiidae (Copepoda: Poecilostomatoida). Biol. Oceanogr. 6(2): 163-173
Huys, R. (1988). Stygofauna of the Canary Islands: 11. Boxshallia bulbantennulata gen. et spec. nov. (Copepoda: Misophrioida) from an anchihaline lava pool on Lanzarote, Canary Islands. Stygologia 4(2): 138-154
Huys, R. (1988). Stygofauna of the Canary Islands: 10. Rotundiclipeidae Fam. Nov. (Copepoda, Harpacticoida) from an anchihaline cave on Tenerife, Canary Islands. Stygologia 4(1): 42-63, more
Huys, R.; Kunz, H. (1988). On the generic boundaries within the marine interstitial Latiremidae (Copepoda: Harpacticoida). Stygologia 4: 292-305, more
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