Bert Hoeksema has been studying Indo-Pacific reef corals since 1982. In that year he was an MSC student and started research on mushroom corals at Naturalis Biodiversity Center, in Leiden, the Netherlands. At present, he is head of the department of Marine Zoology in that institute. Most of his field research was carried out in SE Asia, particularly in Indonesia and Malaysia. He is interested in the taxonomy, phylogeny, biogeography, and ecology of corals and other reef organisms. Much of his present research is focussed on biodiversity patterns, the evolution of inter-specific associations and habitats (phylo-ecology), and on the variability of marine faunas over time (historical ecology).
ERMS 2.0: Costello, M.J.; Bouchet, P.; Boxshall, G.; Arvantidis, C.; Appeltans, W. (2008). European Register of Marine Species. Http://www.marbef.org/data/erms.php. Accessed on yyyy-mm-dd.
[previously: Costello, M.J.; Bouchet, P.; Boxshall, G.; Emblow, C.; Vanden Berghe, E. (2004). European Register of Marine Species. Available online at http://www.marbef.org/data/erms.php. Consulted on [date]., more
Hoeksema, B. W.; Cairns, S. (2024). World List of Scleractinia. Accessed at https://www.marinespecies.org/scleractinia on yyyy-mm-dd, more
Hoeksema, W.; van der Loos, M.; van Moorsel, M. (2022). Coral diversity matches marine park zonation but not economic value of coral reef sites at St. Eustatius, eastern Caribbean. J. Environ. Manage. 320: 115829. https://dx.doi.org/10.1016/j.jenvman.2022.115829
Hoeksema, B.W.; Giyanto; Suharsono (2019). The role of maximum shelf depth versus distance from shore in explaining a diversity gradient of mushroom corals (Fungiidae) off Jakarta. Diversity 11(3): 46. https://dx.doi.org/10.3390/d11030046
Kunihiro, S.; Farenzena, Z.; Hoeksema, B.W.; Groenenberg, D.S.J.; Hermanto, B.; Reimer, J.D. (2019). Morphological and phylogenetic diversity of Waminoa and similar flatworms (Acoelomorpha) in the western Pacific Ocean. Zoology (Jena) 136: 125692. https://dx.doi.org/10.1016/j.zool.2019.06.002
Shelyakin, P.V.; Garushyants, S.K.; Nikitin, M.A.; Mudrova, S.V.; Berumen, M.L.; Speksnijder, A.G.C.L.; Hoeksema, B.W.; Fontaneto, D.; Gelfand, M.S.; Ivanenko, V.N. (2018). Microbiomes of gall-inducing copepod crustaceans from the corals Stylophora pistillata (Scleractinia) and Gorgonia ventalina (Alcyonacea). NPG Scientific Reports 8(1): 10 pp. https://dx.doi.org/10.1038/s41598-018-29953-y
Tarya, A.; Hoitink, A.J.F.; van der Vegt, M.; van Katwijk, M.M.; Hoeksema, B.W.; Bouma, T.J.; Lamers, L.P. M.; Christianen, M.J.A. (2018). Exposure of coastal ecosystems to river plume spreading across a near-equatorial continental shelf. Cont. Shelf Res. 153: 1-15. https://dx.doi.org/10.1016/j.csr.2017.12.003
Valderrama Ballesteros, L.; Matthews, J.L.; Hoeksema, B.W. (2018). Pollution and coral damage caused by derelict fishing gear on coral reefs around Koh Tao, Gulf of Thailand. Mar. Pollut. Bull. 135: 1107-1116. https://dx.doi.org/10.1016/j.marpolbul.2018.08.033
Hoeksema, B.W.; van Beusekom, M.; ten Hove, H.A.; Ivanenko, V.N.; van der Meij, S.E.T.; van Moorsel, G.W.N.M. (2016). Helioseris cucullata as a host coral at St. Eustatius, Dutch Caribbean. Mar. Biodiv. 47(1): 71-78. https://dx.doi.org/10.1007/s12526-016-0599-6
Fransen, C.H.J.M.; Hoeksema, B.W. (2014). Going for the stars: extending the host record for the reef-dwelling Emperor shrimp, Periclimenes imperator (Pontoniinae). Mar. Biodiv. 44(4): 465-466. https://dx.doi.org/10.1007/s12526-014-0210-y
Hoeksema, B.W.; Dekker, F.; de Voogd, N.J. (2014). Free-living mushroom corals strike back by overtopping a coral-killing sponge. Mar. Biodiv. 44(1): 3-4. https://dx.doi.org/10.1007/s12526-013-0188-x
Costello, M.J.; Bouchet, P.; Boxshall, G.; Fauchald, K.; Gordon, D.; Hoeksema, B.W.; Poore, G.C.B.; Van Soest, R.W.M.; Stöhr, S.; Walter, T.C.; Vanhoorne, B.; Decock, W.; Appeltans, W. (2013). Global coordination and standardisation in marine biodiversity through the World Register of Marine Species (WoRMS) and related databases. PLoS One 8(1): 20 pp. https://dx.doi.org/10.1371/journal.pone.0051629
Hoeksema, B.W. (2013). Erratum to: Extreme morphological plasticity enables a free mode of life in Favia gravida at Ascension Island (South Atlantic). Mar. Biodiv. 43(2): 173. http://dx.doi.org/10.1007/s12526-012-0128-1
Santodomingo, N.; Reyes, J.; Flórez, P.; Chacón-Gómez, I.C.; van Ofwegen, L.P.; Hoeksema, B.W. (2013). Diversity and distribution of azooxanthellate corals in the Colombian Caribbean. Mar. Biodiv. 43(1): 7-22. http://dx.doi.org/10.1007/s12526-012-0131-6
van der Meij, S.E.T.; Hoeksema, B.W. (2013). Distribution of gall crabs inhabiting mushroom corals on Semporna reefs, Malaysia. Mar. Biodiv. 43(1): 53-59. https://dx.doi.org/10.1007/s12526-012-0135-2
Waheed, Z.; Hoeksema, B.W. (2013). A tale of two winds: species richness patterns of reef corals around the Semporna peninsula, Malaysia. Mar. Biodiv. 43(1): 37-51. http://dx.doi.org/10.1007/s12526-012-0130-7
Hoeksema, B.W. (2012). Extreme morphological plasticity enables a free mode of life in Favia gravid at Ascension Island (South Atlantic). Mar. Biodiv. 42(2): 289-295. http://dx.doi.org/10.1007/s12526-011-0106-z
Hoeksema, B.W.; Roos, P.J.; Cadée, G.C. (2012). Trans-Atlantic rafting by the brooding reef coral Favia fragum on man-made flotsam. Mar. Ecol. Prog. Ser. 445: 209-218. dx.doi.org/10.3354/meps09460
Hoeksema, B.W.; van der Meij, S.E.T.; Fransen, C.H.J.M. (2012). The mushroom coral as a habitat. J. Mar. Biol. Ass. U.K. 92(4): 647-663. http://dx.doi.org/10.1017/S0025315411001445
Hoeksema, B.W.; Waheed, Z. (2012). It pays to have a big mouth: mushroom corals ingesting salps at northwest Borneo. Mar. Biodiv. 42(2): 297-302. http://dx.doi.org/10.1007/s12526-012-0110-y
Reimer, J.D.; Lin, M.; Fujii, T.; Lane, D.J.W.; Hoeksema, B.W. (2012). The phylogenetic position of the solitary zoanthid genus Sphenopus (Cnidaria: Hexacorallia). Contributions to Zoology 81(1): 34-54
Van Bruggen, A.C.; Hoeksema, B.W. (2012). In memoriam Dr Jacob van der Land (1935-2011), late marine biologist at the Leiden Museum (1964-2000). Zool. Meded. 86: 506-515
Hoeksema, B.W.; Kleemann, K. (2002). New records of Fungiacava eilatensis Goreau et al., 1968 (Bivalvia, Mytilidae) boring into Indonesian mushroom corals (Scleractinia, Fungiidae). Basteria 66: 25-30
Kleemann, K.; Hoeksema, B.W. (2002). Lithophaga (Bivalvia, Mytilidae), including a new species, boring into mushroom corals (Scleractinia, Fungiidae) off South Sulawesi, Indonesia. Basteria 66: 11-24
Hoeksema, B.W. (1991). Evolution of body size in mushroom corals (Scleractinia: Fungiidae) and its ecomorphogical consequences. Neth. J. Zool. 41(2-3): 112-129
Borel Best, M.; Hoeksema, B.W.; Moka, W.; Moll, A.; Suharsono; Nyoman Sutarna, I. (1989). Recent scleractinian coral species collected during the Snellius-II Expedition in eastern Indonesia. Neth. J. Sea Res. 23(2): 107-115. https://dx.doi.org/10.1016/0077-7579(89)90005-7
Hoeksema, B.W. (1989). Taxonomy, phylogeny and biogeography of mushroom corals (Scleractina: Fungiidae). Zoologische Verhandelingen, 254. Rijksmuseum van Natuurlijke Historie Leiden: Leiden. 295, 678 fig. plates 1-6 pp.
Hoeksema, B.W.; Moka, W. (1989). Species assemblages and phenotypes of mushroom corals (Fungiidae) related to coral reef habitats in the Flores Sea. Neth. J. Sea Res. 23(2): 149-160
Hoeksema, B.W. (1981). Het erwtekrabbetje, Pinnotheres pisum (L.), als commensaal van de paardemossel, Modiolus modiolus (L.). Het Zeepaard 41(3): 79
Hoeksema, B.W. (1981). Rugschilden en eiersnoeren van Alloteuthis subulata (Lamarck, 1978). Het Zeepaard 41(5): 144-146
Hoeksema, B.W.; Hoeksema, D. (1979). Naar aanleiding van een Loligo, rugschilden, invasie op Terschelling. Het Zeepaard 39(1): 13-24
Hoeksema, B.W. (1978). Vinstralen in vis en gruis. Het Zeepaard 38(5/6): 116-121
Hoeksema, B.W.; van Ofwegen, L.P. (Ed.) (2004). Fauna Malesiana: Indo-Malayan reef corals: a generic overview [CD-ROM]. World Biodiversity Database CD-ROM Series. Expert Center for Taxonomic Identification (ETI): Amsterdam. ISBN 90-75000-70-7. 1 cd-rom pp.
Gittenberger, A.; Hoeksema, B.W. (2006). Phenotypic plasticity revealed by molecular studies on reef corals of Fungia (Cycloseris) spp. (Scleractinia: Fungiidae) near river outlets, in: Gittenberger, A. The evolutionary history of parasitic gastropods and their coral hostsin the Indo-Pacific = De evolutionaire geschiedenis van parasitaire slakken en hun gastheer koralen in de Indo-Pacific. pp. 28-33
Gittenberger, A.; Hoeksema, B.W. (2006). Habitat preferences of 20 Indo-West Pacific wentletrap species (Gastropoda: Epitoniidae) associated with scleractinian corals, in: Gittenberger, A. The evolutionary history of parasitic gastropods and their coral hostsin the Indo-Pacific = De evolutionaire geschiedenis van parasitaire slakken en hun gastheer koralen in de Indo-Pacific. pp. 217-243
Gittenberger, A.; Reijnen, B.T.; Hoeksema, B.W. (2006). A molecular analysis of the evolutionary history of mushroom corals (Scleractinia: Fungiidae) and its consequences for taxonomic classification, in: Gittenberger, A. The evolutionary history of parasitic gastropods and their coral hostsin the Indo-Pacific = De evolutionaire geschiedenis van parasitaire slakken en hun gastheer koralen in de Indo-Pacific. pp. 37-56
Cairns, S.D.; Hoeksema, B.W.; van der Land, J. (2001). Scleractinia, in: Costello, M.J. et al.European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 109-110, more
Appeltans, W.; Decock, W.; Vanhoorne, B.; Hernandez, F.; Bouchet, P.; Boxshall, G.; Fauchald, K.; Gordon, D.P.; Poore, G.C.B.; Hoeksema, B.W.; van Soest, R.W. (2010). The World Register of Marine Species, the first online inventory of all marine species, in: A decade of discovery: Census of Marine Life 2010. The J. Frederick Grassle Scientific Symposium on the Census of Marine Life: Poster abstracts. pp. [1]
Hoeksema, B.W. (Ed.) (2022). Marine biodiversity of Bonaire: a baseline survey. Preliminary results of the Bonaire Marine Biodiversity Expedition. 2nd Edition. Naturalis Biodiversity Center/ANEMOON Foundation: Leiden, Bennebroek. 78 pp.
Gittenberger, A.; Rensing, M.; Stegenga, H.; Hoeksema, B.W. (2009). Inventarisatie van de aan hard substraat gerelateerde macroflora en macrofauna in de Nederlandse Waddenzee. GiMaRIS Rapport, 2009.11. Marine Research, Inventory and Strategy Solutions: Leiden. 63 pp.
Appeltans, W.; Decock, W.; Vanhoorne, B.; Hernandez, F.; Bouchet, P.; Boxshall, G.; Fauchald, K.; Gordon, D.P.; Hoeksema, B.W.; Poore, G.C.B.; van Soest, R.; Stöhr, S.; Walter, C.; Costello, M.J. (2010). The World Register of Marine Species, the first online inventory of all marine species [Poster]. Vlaams Instituut voor de Zee (VLIZ)/WoRMS Steering Committee: Oostende. 1 poster pp.
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